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Andalusiella punctata
From Fensome et al., 2019:
Palaeocystodinium punctatum Jain and Millepied, 1973, p.29, pl.2, fig.24; pl.3, figs.26–28.
Emendation: Masure et al., 1996, p.180–181, as Andalusiella mauthei subsp. punctata.
Holotype: Jain and Millepied, 1973, pl.3, fig.27.
Originally Palaeocystodinium punctatum, subsequently Andalusiella polymorpha subsp. punctata, thirdly Andalusiella mauthei subsp. punctata (combination illegimate), fourthy (and now) Andalusiella punctata.
Taxonomic senior synonym: Svalbardella (now Andalusiella) polymorpha, according to Lentin and Williams (1976, p.89) — however, Schrank (1987, p.265) retained this taxon as the subspecies Andalusiella polymorpha subsp. punctata.
Taxonomic junior synonym: Andalusiella mauthei, by implication in Masure et al. (1996, p.180), who considered Andaliusiella mauthei (illegitimately) to be the senior name — however, Mahmoud and Schrank (2007, p.604–605) treated "... Andalusiella mauthei as a species excluding the subspecies punctata because the name punctatum [sic] has priority over mauthei and therefore cannot become a subspecies of the latter ...." Mahmoud and Schrank did not indicate how they would alternatively treat "punctata".
Consequently, herein we propose the new combination Andalusiella punctata.
Jain and Millepied (1973, p.32 — figure caption) mislabelled the specimen illustrated in their pl.2, fig.24 as Palaeocystodinium microgranulatum.
Age: Maastrichtian.
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Original description: [Jain and Millepied, 1973]:
Palaeocystodinium punctatum sp. nov.
PI. 2. Fig. 24; PI. 3. Figs. 26-28
Holotype- PI. 3, Fig. 27; SI. No. 8457-5.
Type locality- Bore Hole No. CM-1, Senegal, N-W Africa.
Horizon- Maestrichtian
Diagnosis- Shell spindle-shaped, dorso-ventrally flattened, with two short appendages, one apical and one antapical; Antapical horn divided into another short horn. Archaeopyle trapezoidal below apical horn; periphragm micropunctate. Capsule large, filling completely periphragm cavity, endophragm smooth. Apical horn terminate into a dentate end on free side. No furrows seen. Archaeopyle not observed in all specimens. Another thin membrane exterior to periphragm is present, seen only around horns. The size ratio between horn and central vesicle (H-V ratio) remains mostly 1:4.
Dimensions
Holotype- Shell-length 144 μ, breadth 65 μ; Capsule-length 91 μ, breadth 65 μ; Apical horn-length 22·1 μ, breadth 9·1 μ; Antapical horns-length 31·2 μ, breadth 7·8 μ.
Observed range- Shell-length 140-150 μ, breadth 55-70 μ; Capsule-length 95-97 μ, breadth 60-70 μ; Apical horn-length 20-23 μ, breadth 8-11μ; Antapical horn-length 9-32 μ, breadth 4-8 μ.
Comparison- Palaeocystodinium punctatum sp. novo compares only with P. benjaminii Drugg (1967) in having two antapical horns but differs readily in having no fiber or hairs on the apical horn and having dentate apex, micropunctate periphragm and third exterior layer.
Remarks- The presence of two closely placed antapical horns and horn-vesicle ratio makes it difficult to place this species in Palaeocystodinium and the former evidence might permit the transfer of these forms under a new genus.
Palaeocystodinium punctatum Jain and Millepied, 1973, p.29, pl.2, fig.24; pl.3, figs.26–28.
Emendation: Masure et al., 1996, p.180–181, as Andalusiella mauthei subsp. punctata.
Holotype: Jain and Millepied, 1973, pl.3, fig.27.
Originally Palaeocystodinium punctatum, subsequently Andalusiella polymorpha subsp. punctata, thirdly Andalusiella mauthei subsp. punctata (combination illegimate), fourthy (and now) Andalusiella punctata.
Taxonomic senior synonym: Svalbardella (now Andalusiella) polymorpha, according to Lentin and Williams (1976, p.89) — however, Schrank (1987, p.265) retained this taxon as the subspecies Andalusiella polymorpha subsp. punctata.
Taxonomic junior synonym: Andalusiella mauthei, by implication in Masure et al. (1996, p.180), who considered Andaliusiella mauthei (illegitimately) to be the senior name — however, Mahmoud and Schrank (2007, p.604–605) treated "... Andalusiella mauthei as a species excluding the subspecies punctata because the name punctatum [sic] has priority over mauthei and therefore cannot become a subspecies of the latter ...." Mahmoud and Schrank did not indicate how they would alternatively treat "punctata".
Consequently, herein we propose the new combination Andalusiella punctata.
Jain and Millepied (1973, p.32 — figure caption) mislabelled the specimen illustrated in their pl.2, fig.24 as Palaeocystodinium microgranulatum.
Age: Maastrichtian.
---------------------------------------------------------------------------------------------------
Original description: [Jain and Millepied, 1973]:
Palaeocystodinium punctatum sp. nov.
PI. 2. Fig. 24; PI. 3. Figs. 26-28
Holotype- PI. 3, Fig. 27; SI. No. 8457-5.
Type locality- Bore Hole No. CM-1, Senegal, N-W Africa.
Horizon- Maestrichtian
Diagnosis- Shell spindle-shaped, dorso-ventrally flattened, with two short appendages, one apical and one antapical; Antapical horn divided into another short horn. Archaeopyle trapezoidal below apical horn; periphragm micropunctate. Capsule large, filling completely periphragm cavity, endophragm smooth. Apical horn terminate into a dentate end on free side. No furrows seen. Archaeopyle not observed in all specimens. Another thin membrane exterior to periphragm is present, seen only around horns. The size ratio between horn and central vesicle (H-V ratio) remains mostly 1:4.
Dimensions
Holotype- Shell-length 144 μ, breadth 65 μ; Capsule-length 91 μ, breadth 65 μ; Apical horn-length 22·1 μ, breadth 9·1 μ; Antapical horns-length 31·2 μ, breadth 7·8 μ.
Observed range- Shell-length 140-150 μ, breadth 55-70 μ; Capsule-length 95-97 μ, breadth 60-70 μ; Apical horn-length 20-23 μ, breadth 8-11μ; Antapical horn-length 9-32 μ, breadth 4-8 μ.
Comparison- Palaeocystodinium punctatum sp. novo compares only with P. benjaminii Drugg (1967) in having two antapical horns but differs readily in having no fiber or hairs on the apical horn and having dentate apex, micropunctate periphragm and third exterior layer.
Remarks- The presence of two closely placed antapical horns and horn-vesicle ratio makes it difficult to place this species in Palaeocystodinium and the former evidence might permit the transfer of these forms under a new genus.