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Batioladinium principium
From Fensome et al., 2019:
Batioladinium principium Duxbury, 2018, p.178, pl.9, figs.1–3,7–8,10. Holotype: Duxbury, 2018, pl.9, fig.10. Age: early
Valanginian.
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Original description Duxbury, 2018:
Plate 9, figures 1 – 3, 7, 8, 10
Batioladinium sp. 1 DAVEY 1982, p. 22, pl. 5, figs. 12, 13 only.
Batioladinium sp. 1 Davey 1982 – HEILMANN-CLAUSEN 1987, pl. 2,
figs. 20–22.
Holotype: Plate 9, Figure 10.
Paratype: Plate 9, Figures 1–3.
Type Locality: lower Valanginian, Speeton Clay, Speeton, England. Holotype: Division D3C E.F. Q47.4. Paratype: DivisionD3C E.F. G56.4
Derivation of Name: From the Latin principium, foundation beginning – in reference to the possible origins of both Batioladinium vexillum n. sp. and Batioladinium longi- cornutum (Alberti 1961) Brideaux 1975 from this species.
Diagnosis: A large, elongate pareodinioid dinocyst with a very long, gradually-tapering apical horn and two short, unequal
antapical horns. The left antapical horn is short, but of variable length and slightly extended by irregular surface ornament. The right antapical horn may be absent or represented by a surface bulge and a short, solid projection formed of surface ornament can be present. The archeopyle is apical with the operculum usually fully detached. A deep sulcal notch typical of the genus is invariably present. The surface ornament is granular to irregularly vermiculate and of varying height, largely restricted to the hypocyst and cingular area, with ornament density increasing towards the antapex. Low surface ornament is sometimespresent on the epicyst but not the apical horn; alignment of surface ornament can outline the cingulum.
Dimensions: Holotype (operculum detached): Length – 61 μm.
Width – 35 μm.
Paratype (complete): Length – 147 μm. Width – 35 μm.
Overall (operculum detached): 78 (67) 61 μm × 46 (39) 35 μm.
Specimens Measured: 7.
Remarks: Poulsen (1996, p. 60) included in Batioladinium matyjae Poulsen 1996 all specimens informally named Batioladinium sp. 1 in Davey 1982 (Plate 5, Figures 11 – 13). He designated the specimen illustrated in Davey (op. cit., Plate 5, Figure 11 from the lower Ryazanian of Norfolk) as his paratype. The stratigraphic range of B. matyjae quoted by Poulsen (1996, p. 60), including all of Davey’s specimens of Batioladinium sp. 1, is, “Volgian to Early Valanginian in Britain and Ryazanian (Gochteodinia villosa Zone, Egmontodinium expiratum Subzone) in Denmark”. This is somewhat misleading. As noted by Heilmann-Clausen (1987, p. 22), Davey’s quoted range for Batioladinium sp. 1 (Davey 1982, p. 22) is latest Volgian to early Ryazanian, al-
Batioladinium principium Duxbury, 2018, p.178, pl.9, figs.1–3,7–8,10. Holotype: Duxbury, 2018, pl.9, fig.10. Age: early
Valanginian.
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Original description Duxbury, 2018:
Plate 9, figures 1 – 3, 7, 8, 10
Batioladinium sp. 1 DAVEY 1982, p. 22, pl. 5, figs. 12, 13 only.
Batioladinium sp. 1 Davey 1982 – HEILMANN-CLAUSEN 1987, pl. 2,
figs. 20–22.
Holotype: Plate 9, Figure 10.
Paratype: Plate 9, Figures 1–3.
Type Locality: lower Valanginian, Speeton Clay, Speeton, England. Holotype: Division D3C E.F. Q47.4. Paratype: DivisionD3C E.F. G56.4
Derivation of Name: From the Latin principium, foundation beginning – in reference to the possible origins of both Batioladinium vexillum n. sp. and Batioladinium longi- cornutum (Alberti 1961) Brideaux 1975 from this species.
Diagnosis: A large, elongate pareodinioid dinocyst with a very long, gradually-tapering apical horn and two short, unequal
antapical horns. The left antapical horn is short, but of variable length and slightly extended by irregular surface ornament. The right antapical horn may be absent or represented by a surface bulge and a short, solid projection formed of surface ornament can be present. The archeopyle is apical with the operculum usually fully detached. A deep sulcal notch typical of the genus is invariably present. The surface ornament is granular to irregularly vermiculate and of varying height, largely restricted to the hypocyst and cingular area, with ornament density increasing towards the antapex. Low surface ornament is sometimespresent on the epicyst but not the apical horn; alignment of surface ornament can outline the cingulum.
Dimensions: Holotype (operculum detached): Length – 61 μm.
Width – 35 μm.
Paratype (complete): Length – 147 μm. Width – 35 μm.
Overall (operculum detached): 78 (67) 61 μm × 46 (39) 35 μm.
Specimens Measured: 7.
Remarks: Poulsen (1996, p. 60) included in Batioladinium matyjae Poulsen 1996 all specimens informally named Batioladinium sp. 1 in Davey 1982 (Plate 5, Figures 11 – 13). He designated the specimen illustrated in Davey (op. cit., Plate 5, Figure 11 from the lower Ryazanian of Norfolk) as his paratype. The stratigraphic range of B. matyjae quoted by Poulsen (1996, p. 60), including all of Davey’s specimens of Batioladinium sp. 1, is, “Volgian to Early Valanginian in Britain and Ryazanian (Gochteodinia villosa Zone, Egmontodinium expiratum Subzone) in Denmark”. This is somewhat misleading. As noted by Heilmann-Clausen (1987, p. 22), Davey’s quoted range for Batioladinium sp. 1 (Davey 1982, p. 22) is latest Volgian to early Ryazanian, al-