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Protoellipsodinium crisulatum

From Fensome et al., 2019:
Protoellipsodinium crisulatum

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Original description Duxbury, 2019:

Plate 2, Figures 3–6
Holotype: Plate 2, Figure 3.
Type Locality: Heslerton No. 2 at 11.50 m (core chip), “mid” Aptian. Holotype: E.F. E39.4.

Derivation of Name: From the Latin cristula, a small tuft, comb, plume - in reference to the characteristic crests of this species.

Diagnosis: A smooth, ellipsoidal, thin-walled species with rounded apical and antapical poles. Longitudinal ridges cover the surface of the cyst, and these bear fairly high, serrated crests for much of their length, grading into short, broad-based, acuminate spines at the poles. The archeopyle involves the displacement of a single plate, presumably the mid-dorsal precingular (3'').

Dimensions: Holotype: Length - 56 μm. Width - 41 μm.
Overall: Length - 66 (57) 48 μm; Width - 41 (39) 35 μm
Specimens Measured: 7.

Remarks: Protoellipsodinium cristulatum has longitudinal ridges, with serrated crests over much of its surface, although these grade to broad-based spines at the poles. In having a ridged surface it is most similar to Protoellipsodinium spinocristatum, although that species bears only acuminate spines, which may occasionally coalesce proximally, but without the crests typical of P. cristulatum.

In some respects, P. cristulatum and P. spinocristatum are similar to Ellipsodinium rugulosum Clarke and Verdier 1967, but differ in lacking the clear cingulum and reticulate sculpture specified for that genus by Clarke and Verdier (1967, p. 68). Protoellipsodinium cristulatum and P. spinocristatum also possess crests and/or spines along surface ridges, whereas the muri of E. rugulosum possesses only irregular distal margins (Clarke and Verdier op. cit., p. 69).
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