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Pseudoceratium pelliferum

Pseudoceratium pelliferum Gocht, 1957, p.166–168, pl.18, figs.1–2; text-figs.1–3. Emendation: Dörhöfer and Davies, 1980, p.39.

Holotype: Gocht, 1957, pl.18, fig.1
Locus typicus: Well Ruehlertwist 3, Emsland, NW Germany
Stratum typicum: Valanginian-Hauterivian
Translation Gocht, 1957: LPP

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G.L. Williams short notes on species, Mesozoic-Cenozoic dinocyst course, Urbino, Italy, May 17-22, 1999 - LPP VIEWER CD-ROM 99.5.

Pseudoceratium pelliferum Gocht, 1957, emend. Dörhöfer and Davies, 1980. Diagnosis from Gocht (1957). Cyst flat, in outline asymmetrical-triangular, longer than wide, with strong apical horn and two shorter antapical horns of unequal length. Covered with more or less closely spaced, short, rounded-obtuse, bristle-like processes which are either isolated or are joined with one another at the base. According to Harding (1990b), the length of the three horns is related to water temperature (see Mutterlose and Harding, 1987a, 1987b). The apteate ornamentation may be developed or almost non-existent but is generally densely spaced over the cyst. Size: length 92-173 µm, width 52-70 µm. Holotype: length of apical horn 35 µm, length of longer antapical horn 25 µm, length of shorter antapical horn 12 µm.
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Original description: Gocht 1957, p. 166-168
Diagnosis: Shell flattened, in outline asymmetrically triangular, longer than wide, with strong apical horn and two shorter antapical horns of unequal length. Set more or less densely with short, rounded-blunt, brush-like processes, which stand isolated or are interconnected proximally.
Dimensions: Holotype: Length 132 µm, width 59 µm, apical horn about 35 µm, large antapical horn about 25 µm, small antapical horn about 12 µm. Processes around 2 µm. Other specimens: Length 92-173 (132) µm, width 52-70 (57) µm.

Supplemental description: Pocock, 1962, p. 79
Theca sub-triangular with convex sides; angles prolonged to three horns, relatively long epithecal horn and two shorter, widely divergent, hypothecal horns, one of which may be much reduced, or absent in some specimens; the epithecal region frequently becomes detached along a line of weakness which corresponds to the equatorial girdle, although this structure is not visible in complete specimens. The thecal ornament is variable, comprising spines of varying length arising from a thin membrane closely attached to the theca; the spines are, in general, shorter on the main body of the theca than on the horns; they are usually longest towards the tip of the antapical hypothecal horn where they widen at the tip, bifurcate or, sometimes, branch in a complex manner; along the epithecal horn the spines are well developed in three longitudinal rows which form well marked crests; they are thicker than on other parts of the theca and directed back towards the tip of the spine giving it a serrated outline; the central "crest" of spines running longitudinally down the centre of this spine probably corresponds to the line of plating seen on some species of Ceratium, a related genus. Some specimens of this species show rounded yellow-green organic fragments within the test.
Size Range: Test 75 (89) 95 µm X 69 (77) 85 µm. Hypothecal horns 15 (18) 21 µm. Epithecal horn 45 (47) 51 µm. Spine length 4.5 µm average.

Emended description: Dörhöfer and Davies, 1980, p.39
The species is here emended to account for the presence of two antrior intercalary plates, which are included in the operculum; archaeopyle (4A2I); ornament apteate, short and numerous penitabular and intratabular processes, the latter possibly following latent sutures between fused plates. The surface between the processes finely corrugated-reticulate. Horn positions correspond to one each in apical, left antapical, and right postcingular positions. A slight left postcingular bulge possibly indicates a fourth horn.
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