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Systematophora areolata
Systematophora areolata Klement, 1960
Holotype: Klement, 1960, pl. 9, fig.1-3; Stancliffe and Sarjeant, 1990, pl.5, fig.7
Stratum typicum: Malm
Locus typicus: Well Scherstetten-1, Germany
Translation, Klement 1960, Courtinat 1989: LPP
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G.L. Williams short notes on species, Mesozoic-Cenozoic dinocyst course, Urbino, Italy, May 17-22, 1999 - LPP VIEWER CD-ROM 99.5.
Systematophora areolata Klement, 1960. Systematophora areolata has dorsally 4 smaller epicystal, 3 large hypocystal areolae, ventrally 2 smaller areolae on the epicyst, 4 in groups of two on the hypocyst. The sulcus is made up of 2 elongated narrow fields arranged longitudinally on each surface. The cingulum is composed of 6 isolated narrow, elongate, rectangular fields, each with two processes. Antapex with 1 large, trapezoidal field. Processes relatively long and simple or branched, terminating in sharp point or in a thickening or shortly bifurcate. Size: length 78-109 µm, breadth 72-92 µm, without processes: length 58-63 µm, breadth 46-48 µm. Process length 17-23 µm
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Original description: Klement, 1960, p. 62-64
Diagnosis: typical species of the genus Systematophora, showing the following arrangement of trapezoid areas with convex sides: Dorsal: 4 small epithecal, 3 large hypothecal, always semicircular, parallel to the girdle-region. Ventral: 2 small, oval epithecal, 4 (in 2 groups) similar hypothecal, always symmetrically arouns a longitudinal middle region.
Supplemental description: Courtinat, 1989, p.172
Phragmochorate cyst with a densely granulogemmate wall. The paratabulation is expressed by processes united in annular complexes. The paratabulation is expressed by processes united in ring-complexes. The penitabular processes are attached on a basal crest, which itself is supported by small processes which seem to rise from the ornamentation on the autophragm. The apical, precingular, postcingular and antapical complexes are large. The sulcal processes are grouped in small annular complexes and the cingular processes are isolated and grouped in pairs for every cingular paraplate. The archaeopyle is apical, formed by the loss of 4 apical paraplates. The operculum is simple multiplated. Accessory archaeopyle sutures strongly affect the preapical paraplates. The operculum is also divided by accessory sutures. It is, however, described as a simple rather than a composite archaeopyle, because the accessory sutures are interpreted as being the result of a mechanical action during fossilization. The mechanical ruptures preferably develop along zones of weakness, in this case the paraplate boundaries. The paratabulation formula is, following the Kofoid notation: 4', 0a, 6'', 6c, 6''', 1p, ?s, ?1pv; following the Taylor-Evitt notation: A-C, 1u, 2-6, 1i, au-fi, II-VI, X, Y, ?fu, ?1, ?Z. The archaeopyle type is: (A(1'-4'))/(A(A-C, 1u)).
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Poulsen, 1996, p. 65, Pl. 30, Figs. 4-5
Systematophora areolata Klement 1960
Remarks. The form Systematophora cf. areolata of Davey 1982 is here regarded as S. areolata.
Recorded occurrences.
Denmark: Late Callovian or Early Oxfordian to Ryazanian.
Poland: Transversarium to Scythicus Zones.
Holotype: Klement, 1960, pl. 9, fig.1-3; Stancliffe and Sarjeant, 1990, pl.5, fig.7
Stratum typicum: Malm
Locus typicus: Well Scherstetten-1, Germany
Translation, Klement 1960, Courtinat 1989: LPP
--------------------------------------------------
G.L. Williams short notes on species, Mesozoic-Cenozoic dinocyst course, Urbino, Italy, May 17-22, 1999 - LPP VIEWER CD-ROM 99.5.
Systematophora areolata Klement, 1960. Systematophora areolata has dorsally 4 smaller epicystal, 3 large hypocystal areolae, ventrally 2 smaller areolae on the epicyst, 4 in groups of two on the hypocyst. The sulcus is made up of 2 elongated narrow fields arranged longitudinally on each surface. The cingulum is composed of 6 isolated narrow, elongate, rectangular fields, each with two processes. Antapex with 1 large, trapezoidal field. Processes relatively long and simple or branched, terminating in sharp point or in a thickening or shortly bifurcate. Size: length 78-109 µm, breadth 72-92 µm, without processes: length 58-63 µm, breadth 46-48 µm. Process length 17-23 µm
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Original description: Klement, 1960, p. 62-64
Diagnosis: typical species of the genus Systematophora, showing the following arrangement of trapezoid areas with convex sides: Dorsal: 4 small epithecal, 3 large hypothecal, always semicircular, parallel to the girdle-region. Ventral: 2 small, oval epithecal, 4 (in 2 groups) similar hypothecal, always symmetrically arouns a longitudinal middle region.
Supplemental description: Courtinat, 1989, p.172
Phragmochorate cyst with a densely granulogemmate wall. The paratabulation is expressed by processes united in annular complexes. The paratabulation is expressed by processes united in ring-complexes. The penitabular processes are attached on a basal crest, which itself is supported by small processes which seem to rise from the ornamentation on the autophragm. The apical, precingular, postcingular and antapical complexes are large. The sulcal processes are grouped in small annular complexes and the cingular processes are isolated and grouped in pairs for every cingular paraplate. The archaeopyle is apical, formed by the loss of 4 apical paraplates. The operculum is simple multiplated. Accessory archaeopyle sutures strongly affect the preapical paraplates. The operculum is also divided by accessory sutures. It is, however, described as a simple rather than a composite archaeopyle, because the accessory sutures are interpreted as being the result of a mechanical action during fossilization. The mechanical ruptures preferably develop along zones of weakness, in this case the paraplate boundaries. The paratabulation formula is, following the Kofoid notation: 4', 0a, 6'', 6c, 6''', 1p, ?s, ?1pv; following the Taylor-Evitt notation: A-C, 1u, 2-6, 1i, au-fi, II-VI, X, Y, ?fu, ?1, ?Z. The archaeopyle type is: (A(1'-4'))/(A(A-C, 1u)).
--------------------------------------------
Poulsen, 1996, p. 65, Pl. 30, Figs. 4-5
Systematophora areolata Klement 1960
Remarks. The form Systematophora cf. areolata of Davey 1982 is here regarded as S. areolata.
Recorded occurrences.
Denmark: Late Callovian or Early Oxfordian to Ryazanian.
Poland: Transversarium to Scythicus Zones.