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Taeniophora iunctispina

Taeniophora iunctispina Klement, 1960, p.68–69, pl.10, figs.1–4. Emendation: Sarjeant and Gocht in Sarjeant, 1984a, p.168–169, as Taeniophora iunctispina.

Originally (and now) Taeniophora, subsequently Systematophora. Sarjeant and Gocht in Sarjeant (1984a, p.168) retained this species in Taeniophora.

Holotype: Klement, 1960, pl.10, fig.1-2; Sarjeant, 1984, pl.3, figs.1-2, text-fig.6; Stancliffe and Sarjeant, 1990, pl.3, figs.3,8; Fauconnier and Masure, 2004, pl.78, figs.3–4
Locus typicus: Scherstetten 1 borehole, southwest Germany, at between 1783.1 and 1783.9 m. depth
Stratum typicum: Malm Upper Alpha, middle Oxfordian
Translation Klement, 1960: LPP
Translation Courtinat, 1989: LPP

Original diagnosis: Klement, 1960, p.70
Typical species of the genus Taeniophora with all characteristics and the following special features: processes simple, seldom furcate and distally closed, with elliptical cross-section and widened base developing connecting ridges on the shell surface.

Emended diagnosis: Sarjeant, 1984, p.168
Cyst almost spheroidal, proximate, modified phragmochorate. The length of processes is between 50% and 60% of the cyst breadth. Processes slender or of moderate breadth, distally simple, capitate, bifurcate or trifurcate, the tips of their branches often bifurcating or being extended into trabeculae linking adjacent processes. Lines or low ridges on the phragma surface connect the bases of processes, forming elongate or polygonal structures echoing a penitabulation. At least five linear structures are developed; four or all of these are confluent across the line of the cingulum, which is indicated principally or only by flexures in, or by a waistlike narrowing of, these structures. A sixth, polygonal structure (a peniplate) marks the antapex. Archaeopyle apical, with well-marked sulcal notch and accessory sutures.
Dimensions: Holotype: maximum overall cross-measurement c. 115 µm, length of cyst 46 µm, breadth 58 µm, length of processes up to 35 µm. Range of dimensions; maximum overall cross-measurement c. 100-115 µm, length of cyst 46-52 µm, breadth around 58 µm in all specimens measured, maximum length of processes 29-35 µm.
Description: The surface of the phragma is typically punctate and may also exhibit fine granulation; the processes may show a quite coarse granulation. Four continuous linear process complexes may be seen on the excellently preserved holotype and recognized also, on the paratypes, with variable confidence according to their orientation and condition. Four of these, corresponding probably to paraplates 1" and 2""", 2" and 3""", 3" and 4""", and 4" and 5""", are continuous across the line of the cingulum, which is indicated only by flexures in their margins. The fifth appears to be divided into two sections (peniplates?), joined by a ridge, and apparently corresponds to paraplates 5" and 6""". A polygonal complex corresponds to antapical paraplate 1"""". The processes in each complex are not numerous and appear originally to have been gonal in situation.

Remark:
Courtinat, 1989, p.174: Sarjeant (1984), studying the holotype of Klement, defined this species (and consequently the genus) as being characterized by processes grouped in complexes, these complexes not corresponding to one paraplate like in the genus Systematophora, but to several paraplates. What seems very unusual is that the same complex would reflect one preapical as well as one postapical paraplate. The specimens encountered in the Jura do not strictly present the process arrangement observed by Sarjeant (1984). This arrangement, which is no longer that of Systematophora, is arched or horseshoe-shaped. This characteristic resembles that of the genera Amphorula or Histiophora.
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