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Spiculodinium alatum

From Fensome et al., 2019:

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Original diagnosis Duxbury, 1977, p.49:
A species of Spiniferites with a fairly thick-walled, granular endophragm and periphragm. The periphragm is produced into long (about half main body width) gonal processes which are distally trifurcate. The gonal processes are laterally connected by high, perforate crests which bear bifurcate sutural processes. Archeopyle precingular, probably formed by loss of plate 3".
Observed Dimensions: Holotype 92 x 86 µm. Overall 92(82)72 x 94(76)63 µm. Specimens Measured: 9.

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Remarks Duxbury, 2018:
Spiniferites alatus DUXBURY 1977, p. 49, pl. 5, fig. 4.
Holotype: Spiculodinium alatum (Duxbury 1977, Plate 5, Figure
4) Duxbury n. gen. n. comb.
Remarks: The main difference between this species and
Spiculodinium neptuni is its possession of prominent sutural
crests, linking gonal processes and bifurcate sutural processes
along their length. Otherwise, these species are very similar in
both the wall structure and in the possession of a single plate
(3'') precingular archeopyle. The specimen illustrated in Plate 8,
Figures 4–6 shows the main characteristic features of this species
and clearly illustrates the alignment of processes, marking-
out a gonyaulacacean tabulation, including the cingular and
sulcal areas.
The stratigraphic range of this species was recorded in Duxbury
(1977) as Berriasian to earlyValanginian, but in the current
study it extends into the early Hauterivian.
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