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Chlamydophorella ambigua ssp. hystrix

Plate 35, Figures 1-3, 5, 6, 9
Holotype: Plate 35, Figures 1-3, 9
Type Locality: Speeton Bed C7H, early Hauterivian, inversum ammonite Zone. Holotype: Slide C7H/2, E.F. N54.0.
Derivation of Name: From the Latin hystrix, prickly, thorny – in reference to the minute spines covering each process.

Diagnosis: A small to intermediate, spheroidal cyst with a relatively robust, smooth endocyst bearing numerous short, hollow, proximally and distally flared processes of equal length; each process has a dense cover of minute spines. A delicate ectophragm is supported by the processes. There is no reflected tabulation or apical projection. The archeopyle is apical, with the operculum usually lost.
Dimensions: Holotype: 53 × 58 μm
Overall: 56(49)43 μm × 68(54)46 μm
Specimens Measured: 12

Remarks: The overall morphology of this subspecies is essentially similar to other spheroidal Chlamydophorella spp. that lack an apical projection, including that illustrated in Davey 1978 and transferred to Chlamydophorella huguonioti (Valensi 1955) Davey 1978 primarily because, “The processes are of the stouter type typical of the genus Chlamydophorella”.
Chlamydophorella ambigua subsp. hystrix differs from others in having an ornament of minute spines covering each process. Specimens of Chlamydophorella ambigua subsp. hystrix show some variation of the ectophragm, and the holotype (Plate 35, Figs. 1-3, 9) has isolated processes terminating in discrete pieces of ectophragm, whereas that in Plate 35, Fig. 6 is more continuous.
Occurrences of this subspecies were recorded between Speeton Beds C11B and LB6 (early Hauterivian to early Barremian, regale to variabilis ammonite Zones), and it was particularly common in the late Hauterivian. Offshore, the FAD of C. ambigua subsp. hystrix is associated with the major increase in Hystrichosphaeridium arborispinum defining palynofloral Zone LKP12.
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