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Protoellipsodinium percultum

Plate 1, Figs. 1, 4-6
Holotype: Plate 1, Figure 5, 6
Paratype: Plate 1, Figure 1, 4
Type Locality: Speeton Bed LB2D, early Barremian, fissicostatum ammonite Zone. Holotype: Slide LB2D(5), E.F. G33.3. Paratype: Slide LB2D(1), E.F. F26.1.

Derivation of Name: From the Latin percultus, highly adorned - in reference to the rows of vacuoles running along the spines.

Diagnosis: A smooth, elongate ovoidal to ellipsoidal, robust species with rounded apical and antapical poles which bears numerous long, flexuous, gradually-tapering, acuminate processes. Processes are typically isolated, having circular contacts with the main body, although some proximal flattening and joining of process bases can occur around the archeopyle margin. Each process has a row of small, distinct vacuoles running its full length. Tabulation is indicated only by the single plate precingular archeopyle.

Dimensions: Holotype: Central body - 48 × 35 μm. Overall - 71× 58 μm.
Paratype: Central body - 46 × 38 μm. Overall - 71 × 61μm.
All measured specimens: Central body - 56 (46) 41 μm × 38 (34) 33 μm.
Overall: 73 (64) 53 μm × 61 (53) 46 μm
Specimens Measured: 9.

Remarks: Protoellipsodinium percultum is morphologically similar to Protoellipsodinium spinosum Davey and Verdier 1971, but differs in possessing a row of small, distinct vacuoles running the full length of each process. The FAD of this species is in Speeton Bed LB6, variabilis ammonite Zone, which is also the inception level of P. spinosum, and it was observed to the top of the studied Speeton section here.
In possessing long, flexuous, vacuolate processes, Protoellipsodinium percultum is similar to Protoellipsodinium ornatum Duxbury n. sp. but it is larger, more elongate and has more numerous processes which are distally acuminate, as opposed to the capitate or more complex spiny terminations of P. ornatum.
Vacuoles were also described for Protoellipsodinium vacuolatum Duxbury 2018, but that species is smaller, with short spines and with vacuoles present only at spine bases and running along low crests, where present.
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