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Wanaea digitata
Wanaea digitata Cookson and Eisenack 1958; emend. Woollam 1982
Holotype: Cookson and Eisenack, 1958, pl.9, fig.2
Locus typicus: Wapet's Rough Range well No.1, W Australia
Stratum typicum: Late Jurassic
Translation Courtinat, 1989: LPP
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G.L. Williams short notes on species, Mesozoic-Cenozoic dinocyst course, Urbino, Italy, May 17-22, 1999 - LPP VIEWER CD-ROM 99.5.
Wanaea digitata Cookson and Eisenack, 1958, emend. Woollam, 1982. This species has a relatively narrow paracingular fringe, which is formed from processes that are joined at an equidistant position from the base. Cookson and Eisenack (1958, p58) described it as a narrow fringe composed of finger-like or pointed processes, which, although they may anastomise tangentially or coalesce proximally, remain free distally. Woollam (1983) emended the diagnosis as follows: hypocyst broadly conical with a short rounded antapical horn, epicyst flattened, only slightly apically convex, ellipsoidal in polar view. Archeopyle epicystal, an equatorial line of schism partially separating the simple compound operculum ( the entire epicyst) from the hypocyst. Cyst wall smooth, paratabulation absent, except for a lace-like (posterior paracingular) equatorial flange, which is interrupted along a line corresponding to the area of attachment of the operculum (parasulcus). The equatorial flange consists of radially arranged processes, generally o more than 10 µm in length, usually thin, simple, bifurcate or capitate, consistently linked between one-third to halfway along their length, but free distally. Size: holotype, length 100 µm, width 109 µm. Paratype, width 110 µm, fringe about 9.5 µm.
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Original description: Cookson and Eisenack 1958, p.58
Diagnosis: Shell broadly cone-shaped, narrowing towards a short rounded apex; the edge is ornamented by a narrow fringe composed of finger-like or pointed processes which, although they may anastomose tangentially or coalesce proximally, remain free distally.
Fensome, 1981, p. 55
Synopsis of diagnosis: A speciess of Wanaea with a hemisphaerical to rounded conical hypotract with or without a pronounced antapical horn. the cingular flange is characteristically narrow in width, about one tenth to one twelfth the overall equatorial diameter of the cyst; it is constructed of radial processes that are simple, bifurcate or capitate, linked towards their base and sometimes also elsewhere along their length, but always distally free.
Emended diagnosis: Woollam, 1982, p. 48
Hypocyst broadly conical with a short rounded antapical horn, epicyst flattened, only slightly apically convex, ellipsoidal in polar view. Archeopyle epicystal, an equatorial line of schism partially separating the simple compound operculum (the entire epicyst) from the hypocyst. Cyst wall smooth, paratabulation absent, except for a lace-like (posterior paracingular) equatorial flange, which is interrupted along a line corresponding to the area of attachment of the operculum (parasulcus). The equatorial flange consists of radially arranged processes, generally no more than 10 µm in length, usually thin, simple, bifurcate or capitate, consistently linked between one-third and half way along their length, but free distally.
Remark: The diagnosis is emended to exclude those forms now placed in Wanaea thysanota, which do not accord with the type material of Wanaea digitata.
Courtinat, 1989, p.212: Remarks on Wanaea digitata and W. thysanota:
Fensome (1981) gave evidence for a possible phyletic relation between species of Wanaea and Energlynia. According to this scheme, W. spectabilis, W. digitata, W. thysanota, W. fimbriata and W. clathrata are different ends of the same lineage. Woollam (1982) thinks that such a sequence is difficult to accept because of the spatial and temporal distribution of these species. In the Jura, W. digitata and W. thysanota progressively shade off into one another, considering the ornamentation of the adcingular crests as morphological characteristic. Fensome's scheme does not appear unreal; it may be integrated in the framework of allopatric speciation.
Affinities:
Cookson and Eisenack 1958, p. 58: This species differs from Wanaea spectabilis in the structure of the ornament. In W. spectabilis the processes are united distally so that the edge of the 'lace' is entire, whereas in W.digitata, although the bases of the processes may be united, the edge of the ornament is always frayed.
Fensome, 1981, p. 55: W. digitata is characterized by tyhe structure and narrowness of its cingular flange. The specimen illustrated by Fensome, 1979, has a flange too wide to be typical of this species; this may also be the casee for specimens illustrated by Sarjeant, 1972 and Muir and Sarjeant, 1978.
Holotype: Cookson and Eisenack, 1958, pl.9, fig.2
Locus typicus: Wapet's Rough Range well No.1, W Australia
Stratum typicum: Late Jurassic
Translation Courtinat, 1989: LPP
--------------------------------------------------
G.L. Williams short notes on species, Mesozoic-Cenozoic dinocyst course, Urbino, Italy, May 17-22, 1999 - LPP VIEWER CD-ROM 99.5.
Wanaea digitata Cookson and Eisenack, 1958, emend. Woollam, 1982. This species has a relatively narrow paracingular fringe, which is formed from processes that are joined at an equidistant position from the base. Cookson and Eisenack (1958, p58) described it as a narrow fringe composed of finger-like or pointed processes, which, although they may anastomise tangentially or coalesce proximally, remain free distally. Woollam (1983) emended the diagnosis as follows: hypocyst broadly conical with a short rounded antapical horn, epicyst flattened, only slightly apically convex, ellipsoidal in polar view. Archeopyle epicystal, an equatorial line of schism partially separating the simple compound operculum ( the entire epicyst) from the hypocyst. Cyst wall smooth, paratabulation absent, except for a lace-like (posterior paracingular) equatorial flange, which is interrupted along a line corresponding to the area of attachment of the operculum (parasulcus). The equatorial flange consists of radially arranged processes, generally o more than 10 µm in length, usually thin, simple, bifurcate or capitate, consistently linked between one-third to halfway along their length, but free distally. Size: holotype, length 100 µm, width 109 µm. Paratype, width 110 µm, fringe about 9.5 µm.
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Original description: Cookson and Eisenack 1958, p.58
Diagnosis: Shell broadly cone-shaped, narrowing towards a short rounded apex; the edge is ornamented by a narrow fringe composed of finger-like or pointed processes which, although they may anastomose tangentially or coalesce proximally, remain free distally.
Fensome, 1981, p. 55
Synopsis of diagnosis: A speciess of Wanaea with a hemisphaerical to rounded conical hypotract with or without a pronounced antapical horn. the cingular flange is characteristically narrow in width, about one tenth to one twelfth the overall equatorial diameter of the cyst; it is constructed of radial processes that are simple, bifurcate or capitate, linked towards their base and sometimes also elsewhere along their length, but always distally free.
Emended diagnosis: Woollam, 1982, p. 48
Hypocyst broadly conical with a short rounded antapical horn, epicyst flattened, only slightly apically convex, ellipsoidal in polar view. Archeopyle epicystal, an equatorial line of schism partially separating the simple compound operculum (the entire epicyst) from the hypocyst. Cyst wall smooth, paratabulation absent, except for a lace-like (posterior paracingular) equatorial flange, which is interrupted along a line corresponding to the area of attachment of the operculum (parasulcus). The equatorial flange consists of radially arranged processes, generally no more than 10 µm in length, usually thin, simple, bifurcate or capitate, consistently linked between one-third and half way along their length, but free distally.
Remark: The diagnosis is emended to exclude those forms now placed in Wanaea thysanota, which do not accord with the type material of Wanaea digitata.
Courtinat, 1989, p.212: Remarks on Wanaea digitata and W. thysanota:
Fensome (1981) gave evidence for a possible phyletic relation between species of Wanaea and Energlynia. According to this scheme, W. spectabilis, W. digitata, W. thysanota, W. fimbriata and W. clathrata are different ends of the same lineage. Woollam (1982) thinks that such a sequence is difficult to accept because of the spatial and temporal distribution of these species. In the Jura, W. digitata and W. thysanota progressively shade off into one another, considering the ornamentation of the adcingular crests as morphological characteristic. Fensome's scheme does not appear unreal; it may be integrated in the framework of allopatric speciation.
Affinities:
Cookson and Eisenack 1958, p. 58: This species differs from Wanaea spectabilis in the structure of the ornament. In W. spectabilis the processes are united distally so that the edge of the 'lace' is entire, whereas in W.digitata, although the bases of the processes may be united, the edge of the ornament is always frayed.
Fensome, 1981, p. 55: W. digitata is characterized by tyhe structure and narrowness of its cingular flange. The specimen illustrated by Fensome, 1979, has a flange too wide to be typical of this species; this may also be the casee for specimens illustrated by Sarjeant, 1972 and Muir and Sarjeant, 1978.