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Adnatosphaeridium caulleryi
From Williams, Lentin and Fensome 1998 - Lentin and Williams Index of Fossil Dinoflagellates:
[Adnatosphaeridium caulleryi (Deflandre, 1939a, p. 189, pl.11, figs.2-3) Williams and Downie, 1969, p. 17; Emendation: Stancliffe and Sarjeant, 1990, p. 200
Originally Hystrichosphaeridium, subsequently Cannosphaeropsis, thirdly (and now) Adnatosphaeridium, fourthly Polystephanephorus. Stancliffe and Sarjeant (1990, p.200) retained this species in Adnatosphaeridium. This combination was not validly published in Williams and Downie (1966c, p.218), since these authors did not fully reference the basionym.
Holotype: Deflandre, 1939a, pl.11, fig.2; Stancliffe and Sarjeant, 1990, pl.1, figs.3,5.]
Locus typicus: Villers-sur-Mer, Calvados, France
Age: Early Oxfordian
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Emended description: [Stancliffe and Sarjeant, 1990, p.200]:
Diagnosis:
Skolochorale dinoflagellate cysts, with central body spherical to subspherical and of intermediate to large overall size. Phragma smooth to finely granulate. Central body surmounted by numerous solid, slender processes which are consistently bifurcate or multifurcate and may be distally expanded. Processes may be deeply or medially furcate and occasionally branch proximally. Processes variably interconnected distally by solid, slender, smooth to irregularly denticulate trabeculae.
Paratabulation gonyaulacacean, indicated by intratabular processes (or clusters of processes). Process clusters irregular, occasionally exhibiting annulate or arcuate organisation: this is shown best in the precingular, postcingular and antapical areas. Apical, cingular, sulcal and posterior intercalary paraplates give rise to single processes or to small, closely spaced process clusters. Trabeculae may or may not interconnect process clusters.
Paratabulation formula probably 4', 6". 6c, 6''', 1p, 1'''', x- ?s.
Archeopyle apical, type (tA), operculum free.
Description:
This species of Adnatosphaeridium has a spherical to subspherical, smooth to finely granulate cyst body bearing numerous thin, solid processes, interpreted as extensions of the periphragm. The process length is variable within a population (20-31% of the overall cyst diameter), but relatively consistent on any individual specimen. The processes, though consistently arcate, are otherwise variable in distal morphology: they may be expanded and interconnected by irregularly denticulate trabeculae. Medial branching of the processes may occur and rarely they bifumate proximally. The lager paraplate areas (precingular, postcingular and antapical) are characterised by distinct intratabular process clusters which are typically irregular, rarely annulate or arcuate, in organisation. Individual processes within these relatively large clusters are normally trabeculate. The size, complexity and trabecular connections between adjacent clusters render some precingular and postcingular clusters difficult to distinguish. The smaller apical, paracingular, parasulcal and posterior intercalary paraplates have solitary, generally bifurcate, processes or small, closely paced process clusters. The large number and complexity of he processes renders the paratabulation formula difficult to determine precisely; however, it appears to be standard gonyaulacacean.
The archaeopyle is apical, type (tA), with a free operculum.
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Notes:
G.L. Williams short notes on species, Mesozoic-Cenozoic dinocyst course, Urbino, Italy, May 17-22, 1999 - LPP VIEWER CD-ROM 99.5.
Adnatosphaeridium caulleryi (Deflandre, 1939a) Williams and Downie, 1969, emend. Stancliffe and Sarjeant, 1990. According to Stancliffe and Sarjeant (1990, p.220)), has numerous solid, slender processes which are consistently bifurcate or multifurcate and may be distally expanded. Processes may be branched, variably interconnected distally by solid, slender, smooth to irregularly denticulate trabeculae. Process clusters irregular, occasionally forming annulate or arcuate complexes, especially in precingular, postcingular and antapical regions. Apical, cingular, sulcal and posterior intercalary plates have single processes. Trabeculae may or connect process clusters.
[Adnatosphaeridium caulleryi (Deflandre, 1939a, p. 189, pl.11, figs.2-3) Williams and Downie, 1969, p. 17; Emendation: Stancliffe and Sarjeant, 1990, p. 200
Originally Hystrichosphaeridium, subsequently Cannosphaeropsis, thirdly (and now) Adnatosphaeridium, fourthly Polystephanephorus. Stancliffe and Sarjeant (1990, p.200) retained this species in Adnatosphaeridium. This combination was not validly published in Williams and Downie (1966c, p.218), since these authors did not fully reference the basionym.
Holotype: Deflandre, 1939a, pl.11, fig.2; Stancliffe and Sarjeant, 1990, pl.1, figs.3,5.]
Locus typicus: Villers-sur-Mer, Calvados, France
Age: Early Oxfordian
-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
Emended description: [Stancliffe and Sarjeant, 1990, p.200]:
Diagnosis:
Skolochorale dinoflagellate cysts, with central body spherical to subspherical and of intermediate to large overall size. Phragma smooth to finely granulate. Central body surmounted by numerous solid, slender processes which are consistently bifurcate or multifurcate and may be distally expanded. Processes may be deeply or medially furcate and occasionally branch proximally. Processes variably interconnected distally by solid, slender, smooth to irregularly denticulate trabeculae.
Paratabulation gonyaulacacean, indicated by intratabular processes (or clusters of processes). Process clusters irregular, occasionally exhibiting annulate or arcuate organisation: this is shown best in the precingular, postcingular and antapical areas. Apical, cingular, sulcal and posterior intercalary paraplates give rise to single processes or to small, closely spaced process clusters. Trabeculae may or may not interconnect process clusters.
Paratabulation formula probably 4', 6". 6c, 6''', 1p, 1'''', x- ?s.
Archeopyle apical, type (tA), operculum free.
Description:
This species of Adnatosphaeridium has a spherical to subspherical, smooth to finely granulate cyst body bearing numerous thin, solid processes, interpreted as extensions of the periphragm. The process length is variable within a population (20-31% of the overall cyst diameter), but relatively consistent on any individual specimen. The processes, though consistently arcate, are otherwise variable in distal morphology: they may be expanded and interconnected by irregularly denticulate trabeculae. Medial branching of the processes may occur and rarely they bifumate proximally. The lager paraplate areas (precingular, postcingular and antapical) are characterised by distinct intratabular process clusters which are typically irregular, rarely annulate or arcuate, in organisation. Individual processes within these relatively large clusters are normally trabeculate. The size, complexity and trabecular connections between adjacent clusters render some precingular and postcingular clusters difficult to distinguish. The smaller apical, paracingular, parasulcal and posterior intercalary paraplates have solitary, generally bifurcate, processes or small, closely paced process clusters. The large number and complexity of he processes renders the paratabulation formula difficult to determine precisely; however, it appears to be standard gonyaulacacean.
The archaeopyle is apical, type (tA), with a free operculum.
-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
Notes:
G.L. Williams short notes on species, Mesozoic-Cenozoic dinocyst course, Urbino, Italy, May 17-22, 1999 - LPP VIEWER CD-ROM 99.5.
Adnatosphaeridium caulleryi (Deflandre, 1939a) Williams and Downie, 1969, emend. Stancliffe and Sarjeant, 1990. According to Stancliffe and Sarjeant (1990, p.220)), has numerous solid, slender processes which are consistently bifurcate or multifurcate and may be distally expanded. Processes may be branched, variably interconnected distally by solid, slender, smooth to irregularly denticulate trabeculae. Process clusters irregular, occasionally forming annulate or arcuate complexes, especially in precingular, postcingular and antapical regions. Apical, cingular, sulcal and posterior intercalary plates have single processes. Trabeculae may or connect process clusters.