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Reutlingia gochtii
Reutlingia gochtii Drugg, 1978, p.72–73, pl.7, figs.7–10. Emendation: Below, 1987a, p.137–138.
Holotype: Drugg, 1978, pl. 7, fig.8-10
Isotype: Drugg, 1978, pl.7, fig.7
Locus typicus: Reutlingen, Germany
Stratum typicum: Callovian
Translation Below, 1987: LPP
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G.L. Williams short notes on species, Mesozoic-Cenozoic dinocyst course, Urbino, Italy, May 17-22, 1999 - LPP VIEWER CD-ROM 99.5.
Reutlingia gochtii Drugg, 1978, emend. Below, 1987a. R. gochtii is ovoidal to rectangular with broad paracingulum (about 8 µm wide) and delineated by low broad based, acuminate projections. Surface of cyst ornamented with broad based, acuminate projections 4-6 µm in height and which are open to the interior. At times they are connected proximally as clusters and tend to be arranged as simulate complexes. Autophragm thin and smooth to delicately rugulate. Length: 26-50 µm. Archeopyle type I attached. Note emendation by Below (1987a). Archeopyle anterior intercalary resulting from loss of 2a, which is adnate.
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Original description: Drugg 1978, p. 72
The cyst is ovoidal to somewhat rectangular in outline. The paracingulum is broad (about 8 µm wide) and deeply incised. It is devoid of ornamentation and is delineated by low, broad based, acuminate projections. The parasulcus is less well delineated but is still fairly distinct. It broadens antapically. The surface of the cyst is ornamented by broad based acuminate projections from 4 to 6 µm in length and which are open into the interior. At times they are connected proximally as clusters and they tend to be arranged as simulate complexes although this feature is not always clearly expressed. the paratabulation is probably 1', 5'', 0c, 5''', 1''''. The archaeopyle is apparently apical o the extent that one specimen was recovered with the apical area missing. All other specimens were complete with no indication of any rupture. The autophragm is thin and smooth to delicately rugulate. The total length ranges from 26 to 50 µm and the total width ranges from 28 to 46 µm. Most specimens are about 34 µm in length and 32 µm in width. The ornamentation is usually about 4 µm in height. Dimensions and description based on 14 specimens.
Emended description: Below, 1987, p. 137-138
Vesicle-/plate arrangement of the dinoflagellate amphiesma cop, pop, cap, 5', 3a,
7''-6'', 7c, 5''', 2'''', as, y, z, ps. 1'''' and 2'''' symmetrically on both sides of the dorso-ventral plane. Large ps omegaform. Growth of thecal plates peridinoid.
Cyst proximate, acavate, cornucavate, apiculocavate, peridinoidal with flattened poles, surface smooth or intratabularly bulged, lenticular, reniform, epicyst and hypocyst of equal width, separated by a very broad, planar to laevorotatory, deeply indented cingulum. Sulcus flat to deeply indented. Small cyst. Wall consisting of thin pedium and luxuria which forms high, hollow, broad-based, truncate, distally oblate, capitate, cauliflorate processes. Arrangement in penitabular rows, in gonal, intergonal and rarely intratabular positions. Processes often connected proximally, building penitabular and also limbate pericoels. Luxuria bordering pedium in intratabular, cingular and sulcal, mostly also limbate, positions, but also tightened tent-like over the plate between the penitabular projections (platepericoel), thus forming marked tabular bulges on the cyst surface; luxurua missing near 2c and 6c. Surface smooth, microfrangate. Tabulation XPR, 5', 3a, 7'', Xc, 5''', 2'''', Xs. cop very small, round, in the form of a peg or ring, horse-shoe-shaped pop and small rhombic cap between 1'/5' and pop. 5 apicals: VI 1' (V), large VI 2' (VL), deltaform inverse- camerate 3' (DL) and 4' (DR), large VIII 5' (VR). 3 anterior intercalaries: linteloid 1a (LDL), Vi or VII 2a (D), inverse-camerate 3a (DR). 7 precingulars: anteriorly geniculate V-nE 1'' (VVL), V-nE 2'' (LVL), V-nE 5'' (DDR) and V-nE 6''(R), anteriorly linear 3'' (LDL), anteriorly geniculate V-nE 4'' (DDL) or anteriorly linear IV-nE 4'' in case of heptagonal 2a, anteriorly linear IV-nE 7'' (VVR), rarely connected with 1'. Cingulum nontabular, leavorotatory. 5 postcingulars: V-nE 1''' (VVL), posteriorly linear IV-nE 2''' (L), IV-nE 4''' (DR) and IV-nE 5''' (VR), posteriorly geniculate V-nE 3''' (DDL), finis 3'''/''' dorsal.
Large V 1'''' (L) and large V 2'''' on both sides of the apex. Sulcus nontabular. Archaeopyle anterior intercalary, 2a. operculum monotabular, adnate at the precingulars, opercular formula 2a(a).
Affinities:
Drugg, 1978, p. 72-73
The genus Stenopyxinium Deflandre, 1968, is similar to the extent that it possesses a deeply incised paracingulum. The ornamentation and, to a lesser extent, the mode of archaeopyle formation appear different. In addition Stenopyxinium shows no sign of paratabulation aside from the archaeopyle and paracingulum. the closest comparison is with Horologinella lineata Cookson and Eisenack, 1962, and H. spinosigibberosa Brideaux & Fisher, 1976. They are similar in outline, exhibit paratabulation, and seemingly possess apical archaeopyles. The ornamentation and wall structure are dissimilar to that of reutlingia. The exact paratabultion is questionable as is that of Reutlingia but they do not seem to be identical.
Below, 1987, p. 138
R. gochtii differs from R. cardobarbata in that the processes are coarser and less numerous. Between the gonal processes there are only 1 or 2 processes, or they are absent. The plates are generally devoid of ornaments. Furthermore, the outline of the cyst in dorso-ventral view is biconical to almost rectangular. Plate 1'''' is built like all other plates, and not covered by a tectum, as it is the case in R. cardobarbata, R. hirsuta or R. fausta. In R. gochtii, the luxuria is mostly missing in the positions of 2c and 6c, so that the chagrinate, granulate pedium-surface is visible. Like in R. cardobarbata and R. fausta, reduction of the number of precingulars to 6 instead of the normally typical 7 is also observed in R. gochtii.
Holotype: Drugg, 1978, pl. 7, fig.8-10
Isotype: Drugg, 1978, pl.7, fig.7
Locus typicus: Reutlingen, Germany
Stratum typicum: Callovian
Translation Below, 1987: LPP
--------------------------------------------------
G.L. Williams short notes on species, Mesozoic-Cenozoic dinocyst course, Urbino, Italy, May 17-22, 1999 - LPP VIEWER CD-ROM 99.5.
Reutlingia gochtii Drugg, 1978, emend. Below, 1987a. R. gochtii is ovoidal to rectangular with broad paracingulum (about 8 µm wide) and delineated by low broad based, acuminate projections. Surface of cyst ornamented with broad based, acuminate projections 4-6 µm in height and which are open to the interior. At times they are connected proximally as clusters and tend to be arranged as simulate complexes. Autophragm thin and smooth to delicately rugulate. Length: 26-50 µm. Archeopyle type I attached. Note emendation by Below (1987a). Archeopyle anterior intercalary resulting from loss of 2a, which is adnate.
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Original description: Drugg 1978, p. 72
The cyst is ovoidal to somewhat rectangular in outline. The paracingulum is broad (about 8 µm wide) and deeply incised. It is devoid of ornamentation and is delineated by low, broad based, acuminate projections. The parasulcus is less well delineated but is still fairly distinct. It broadens antapically. The surface of the cyst is ornamented by broad based acuminate projections from 4 to 6 µm in length and which are open into the interior. At times they are connected proximally as clusters and they tend to be arranged as simulate complexes although this feature is not always clearly expressed. the paratabulation is probably 1', 5'', 0c, 5''', 1''''. The archaeopyle is apparently apical o the extent that one specimen was recovered with the apical area missing. All other specimens were complete with no indication of any rupture. The autophragm is thin and smooth to delicately rugulate. The total length ranges from 26 to 50 µm and the total width ranges from 28 to 46 µm. Most specimens are about 34 µm in length and 32 µm in width. The ornamentation is usually about 4 µm in height. Dimensions and description based on 14 specimens.
Emended description: Below, 1987, p. 137-138
Vesicle-/plate arrangement of the dinoflagellate amphiesma cop, pop, cap, 5', 3a,
7''-6'', 7c, 5''', 2'''', as, y, z, ps. 1'''' and 2'''' symmetrically on both sides of the dorso-ventral plane. Large ps omegaform. Growth of thecal plates peridinoid.
Cyst proximate, acavate, cornucavate, apiculocavate, peridinoidal with flattened poles, surface smooth or intratabularly bulged, lenticular, reniform, epicyst and hypocyst of equal width, separated by a very broad, planar to laevorotatory, deeply indented cingulum. Sulcus flat to deeply indented. Small cyst. Wall consisting of thin pedium and luxuria which forms high, hollow, broad-based, truncate, distally oblate, capitate, cauliflorate processes. Arrangement in penitabular rows, in gonal, intergonal and rarely intratabular positions. Processes often connected proximally, building penitabular and also limbate pericoels. Luxuria bordering pedium in intratabular, cingular and sulcal, mostly also limbate, positions, but also tightened tent-like over the plate between the penitabular projections (platepericoel), thus forming marked tabular bulges on the cyst surface; luxurua missing near 2c and 6c. Surface smooth, microfrangate. Tabulation XPR, 5', 3a, 7'', Xc, 5''', 2'''', Xs. cop very small, round, in the form of a peg or ring, horse-shoe-shaped pop and small rhombic cap between 1'/5' and pop. 5 apicals: VI 1' (V), large VI 2' (VL), deltaform inverse- camerate 3' (DL) and 4' (DR), large VIII 5' (VR). 3 anterior intercalaries: linteloid 1a (LDL), Vi or VII 2a (D), inverse-camerate 3a (DR). 7 precingulars: anteriorly geniculate V-nE 1'' (VVL), V-nE 2'' (LVL), V-nE 5'' (DDR) and V-nE 6''(R), anteriorly linear 3'' (LDL), anteriorly geniculate V-nE 4'' (DDL) or anteriorly linear IV-nE 4'' in case of heptagonal 2a, anteriorly linear IV-nE 7'' (VVR), rarely connected with 1'. Cingulum nontabular, leavorotatory. 5 postcingulars: V-nE 1''' (VVL), posteriorly linear IV-nE 2''' (L), IV-nE 4''' (DR) and IV-nE 5''' (VR), posteriorly geniculate V-nE 3''' (DDL), finis 3'''/''' dorsal.
Large V 1'''' (L) and large V 2'''' on both sides of the apex. Sulcus nontabular. Archaeopyle anterior intercalary, 2a. operculum monotabular, adnate at the precingulars, opercular formula 2a(a).
Affinities:
Drugg, 1978, p. 72-73
The genus Stenopyxinium Deflandre, 1968, is similar to the extent that it possesses a deeply incised paracingulum. The ornamentation and, to a lesser extent, the mode of archaeopyle formation appear different. In addition Stenopyxinium shows no sign of paratabulation aside from the archaeopyle and paracingulum. the closest comparison is with Horologinella lineata Cookson and Eisenack, 1962, and H. spinosigibberosa Brideaux & Fisher, 1976. They are similar in outline, exhibit paratabulation, and seemingly possess apical archaeopyles. The ornamentation and wall structure are dissimilar to that of reutlingia. The exact paratabultion is questionable as is that of Reutlingia but they do not seem to be identical.
Below, 1987, p. 138
R. gochtii differs from R. cardobarbata in that the processes are coarser and less numerous. Between the gonal processes there are only 1 or 2 processes, or they are absent. The plates are generally devoid of ornaments. Furthermore, the outline of the cyst in dorso-ventral view is biconical to almost rectangular. Plate 1'''' is built like all other plates, and not covered by a tectum, as it is the case in R. cardobarbata, R. hirsuta or R. fausta. In R. gochtii, the luxuria is mostly missing in the positions of 2c and 6c, so that the chagrinate, granulate pedium-surface is visible. Like in R. cardobarbata and R. fausta, reduction of the number of precingulars to 6 instead of the normally typical 7 is also observed in R. gochtii.