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Wanneria listeri
From Fensome et al., 2019:
Wanneria listeri (Stover and Helby, 1987a, p.121–122,124, figs.21A–C,22A–D,23A–L) Below, 1987a, p.77,80.
Emendation: Below, 1987a, p.77,80, as Wanneria listeri.
Holotype: Stover and Helby, 1987a, figs.22A–D; Fensome et al., 1996, figs.1–4 — p.2195.
Originally Suessia, subsequently (and now) Wanneria. Age: Norian.
Paratypes: Stover and Helby, 1987, fig. 23A-J
Locus typicus: Unnamed interval (2419.1 m) of Well Eendracht-1, Carnarvon Basin, Western Australia
Stratum typicum: Norian
Translation Below, 1987: Geological Survey of Canada
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Original description: Stover and Helby, 1987, p. 121-124: Suessia listeri
(annotated): Cysts proximate and subspherical. Autophragm smooth, thin (about 1 µm or less) surface divided into numerous, polygonal paraplates delimited by parasutural ridges with distinctly spinate crests. Paraplates 4- to 7-sided (the majority ranging in maximum dimension from 6-15 µm, (most are in the 10-12 µm range), and having narrow relatively low bounding ridges (their width and height are less than 2.5 µm). Crestal spines nearly always solid, some connected proximally, tapered distally, with acuminate, or blunt or less frequently bifurcate tips. Spines 1.5-5 µm long (generally about 3 µm), and their length fairly uniform on individual specimens. Specimens without opercula having 56-64 paraplates; those with opercula have an additional 12-19 paraplates. In the equatorial and adjacent areas paraplates arranged in 3, fairly distinct, latitudinal rows; elsewhere their apparent latitudinal alignment less evident. Paratabulation 0-2pr, 4-6', 6-10a, 0-?2ap, 12-14'', 12c, 12''', 11-16p, 5-6pa, 1'''', and 3s. Combination archeopyle, type [tAtI], usually formed by the release of all paraplates anterior to the precingular series. On occasional specimens, extremities of principal archeopyle suture not meeting, and operculum remaining adherent. Position of the parasulcus inferred by offsetting of cingular and postcingular series and by generally incompletely outlined paraplates (up to 3) on the hypocyst in the area of the offset. Specimens 52(63)80 µm in width; those without opercula 40(51)60 µm in length; specimens with opercula 14 to 16 µm longer; 25 specimens measured.
Emended description: Below, 1987, p. 77-81: Wanneria listeri
Prognosticated arrangement of vesicles/plates on the dinoflagellate amphiesma cop, pop, cap, 5', 3a-4a, 7'', 7c, 5'''-6''', 2'''', ns.
Cystic habit proximochorate, spheroidal, size large; wall composed of pedium with only finate luxuria; surface glabrous, divided into areae by finate rows of spines, the spines expanding basally and standing isolated or connected by basal crests, distally tapered and acute or thickened or bifurcate, the bifurcation perpendicular to the finis so that each of the two tips points to the neighbouring area, short spines often fused together over a variable portion of their height; inner surface polygonally areate, smooth, with the fines in the form of grooves; areation multiareate, multiserial, areation formula n4, n3, n2, n1, nc, n1, n2, n3, n4, sas, pas, rs, ls, pps, sps; areae vary individually in shape and number, ca. 45-90 small tetragonal to octagonal areae distributed as follows: apical quartanterior series n4(1-2)* (* = numbers exemplified in the present material), multiareate tertanterior n3(2-6)* , secanterior n2 (6-9)*, primanterior n1 (10-13)*, multiareate cingulum c(9-15)*, multiareate primposterior series n1(10-15)*, secposterior n2 (6-12)*, tertposterior n3 (1-2)* as well as intercalary areae in variable positions; cingulum equatorial, cingulum and area series helicoid laevorotatory and displaced at the sulcus, beginning with small 1c; sulcus subdivided into sas, pas, rs, ls, pps, and sps, archaeopyle anterior to the primanterior series, tn4 + tn3 + tn2, operculum foederate, secate, opercular formula (tn4 + tn3 + tn2)s.
Affinities:
Stover and Helby, 1987, p. 122, 124: Suessia listeri
Suessia listeri differs from S. swabiana Morbey 1975 in having spinate crests on the parasutural ridges, in being wider and in details of paratabulation. Because of uncertainties about the number of paraplates in each species, differences in the totals are not regarded as critical. However, identification of a postapical series in S. swabiana comprising 7-9 paraplates, compared to the apparent absence of the series in S. listeri, is considered significant. However, 2 opercular paraplates (designated as 13 and 14 on Fig. 21c might be interpreted as postapical paraplates.
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G.L. Williams short notes on species, Mesozoic-Cenozoic dinocyst course, Urbino, Italy, May 17-22, 1999 - LPP VIEWER CD-ROM 99.5.
Wanneria listeri (Stover and Helby, 1987a) Below, 1987a, emend. Below, 1987a. Wanneria listeri differs from Suessia swabiana in having spinate crests on the parasutural ridges, in being wider and in details of tabulation. The tabulation formula is 0-2pr, 4-6', 6-10a, 0-?2ap, 12-14", 12c, 12"’, 11-16p, 5-6pa, 1"”. In a comparison with Suessia swabiana, Stover and Helby (1987, p.122) give the plate formula for Wanneria listeri as: no preapical, ?4 apical, ?3-6 anterior intercalary, 7-9 postapical, 12-14 precingular, 12-14 cingular, 12-14 postcingular,7-9 posterior intercalary, 6 preantapical, 1 antapical, 4-8 sulcal. Archeopyle combination type, usually formed by the release of all paraplates anterior to the precingular series. Below (1987a, p.77,80) emended the diagnosis of this species. Size: length without operculum 40-60 µm, with opercula 14-16 µm longer, width 52-80 µm.
Wanneria listeri (Stover and Helby, 1987a, p.121–122,124, figs.21A–C,22A–D,23A–L) Below, 1987a, p.77,80.
Emendation: Below, 1987a, p.77,80, as Wanneria listeri.
Holotype: Stover and Helby, 1987a, figs.22A–D; Fensome et al., 1996, figs.1–4 — p.2195.
Originally Suessia, subsequently (and now) Wanneria. Age: Norian.
Paratypes: Stover and Helby, 1987, fig. 23A-J
Locus typicus: Unnamed interval (2419.1 m) of Well Eendracht-1, Carnarvon Basin, Western Australia
Stratum typicum: Norian
Translation Below, 1987: Geological Survey of Canada
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Original description: Stover and Helby, 1987, p. 121-124: Suessia listeri
(annotated): Cysts proximate and subspherical. Autophragm smooth, thin (about 1 µm or less) surface divided into numerous, polygonal paraplates delimited by parasutural ridges with distinctly spinate crests. Paraplates 4- to 7-sided (the majority ranging in maximum dimension from 6-15 µm, (most are in the 10-12 µm range), and having narrow relatively low bounding ridges (their width and height are less than 2.5 µm). Crestal spines nearly always solid, some connected proximally, tapered distally, with acuminate, or blunt or less frequently bifurcate tips. Spines 1.5-5 µm long (generally about 3 µm), and their length fairly uniform on individual specimens. Specimens without opercula having 56-64 paraplates; those with opercula have an additional 12-19 paraplates. In the equatorial and adjacent areas paraplates arranged in 3, fairly distinct, latitudinal rows; elsewhere their apparent latitudinal alignment less evident. Paratabulation 0-2pr, 4-6', 6-10a, 0-?2ap, 12-14'', 12c, 12''', 11-16p, 5-6pa, 1'''', and 3s. Combination archeopyle, type [tAtI], usually formed by the release of all paraplates anterior to the precingular series. On occasional specimens, extremities of principal archeopyle suture not meeting, and operculum remaining adherent. Position of the parasulcus inferred by offsetting of cingular and postcingular series and by generally incompletely outlined paraplates (up to 3) on the hypocyst in the area of the offset. Specimens 52(63)80 µm in width; those without opercula 40(51)60 µm in length; specimens with opercula 14 to 16 µm longer; 25 specimens measured.
Emended description: Below, 1987, p. 77-81: Wanneria listeri
Prognosticated arrangement of vesicles/plates on the dinoflagellate amphiesma cop, pop, cap, 5', 3a-4a, 7'', 7c, 5'''-6''', 2'''', ns.
Cystic habit proximochorate, spheroidal, size large; wall composed of pedium with only finate luxuria; surface glabrous, divided into areae by finate rows of spines, the spines expanding basally and standing isolated or connected by basal crests, distally tapered and acute or thickened or bifurcate, the bifurcation perpendicular to the finis so that each of the two tips points to the neighbouring area, short spines often fused together over a variable portion of their height; inner surface polygonally areate, smooth, with the fines in the form of grooves; areation multiareate, multiserial, areation formula n4, n3, n2, n1, nc, n1, n2, n3, n4, sas, pas, rs, ls, pps, sps; areae vary individually in shape and number, ca. 45-90 small tetragonal to octagonal areae distributed as follows: apical quartanterior series n4(1-2)* (* = numbers exemplified in the present material), multiareate tertanterior n3(2-6)* , secanterior n2 (6-9)*, primanterior n1 (10-13)*, multiareate cingulum c(9-15)*, multiareate primposterior series n1(10-15)*, secposterior n2 (6-12)*, tertposterior n3 (1-2)* as well as intercalary areae in variable positions; cingulum equatorial, cingulum and area series helicoid laevorotatory and displaced at the sulcus, beginning with small 1c; sulcus subdivided into sas, pas, rs, ls, pps, and sps, archaeopyle anterior to the primanterior series, tn4 + tn3 + tn2, operculum foederate, secate, opercular formula (tn4 + tn3 + tn2)s.
Affinities:
Stover and Helby, 1987, p. 122, 124: Suessia listeri
Suessia listeri differs from S. swabiana Morbey 1975 in having spinate crests on the parasutural ridges, in being wider and in details of paratabulation. Because of uncertainties about the number of paraplates in each species, differences in the totals are not regarded as critical. However, identification of a postapical series in S. swabiana comprising 7-9 paraplates, compared to the apparent absence of the series in S. listeri, is considered significant. However, 2 opercular paraplates (designated as 13 and 14 on Fig. 21c might be interpreted as postapical paraplates.
--------------------------------------------------
G.L. Williams short notes on species, Mesozoic-Cenozoic dinocyst course, Urbino, Italy, May 17-22, 1999 - LPP VIEWER CD-ROM 99.5.
Wanneria listeri (Stover and Helby, 1987a) Below, 1987a, emend. Below, 1987a. Wanneria listeri differs from Suessia swabiana in having spinate crests on the parasutural ridges, in being wider and in details of tabulation. The tabulation formula is 0-2pr, 4-6', 6-10a, 0-?2ap, 12-14", 12c, 12"’, 11-16p, 5-6pa, 1"”. In a comparison with Suessia swabiana, Stover and Helby (1987, p.122) give the plate formula for Wanneria listeri as: no preapical, ?4 apical, ?3-6 anterior intercalary, 7-9 postapical, 12-14 precingular, 12-14 cingular, 12-14 postcingular,7-9 posterior intercalary, 6 preantapical, 1 antapical, 4-8 sulcal. Archeopyle combination type, usually formed by the release of all paraplates anterior to the precingular series. Below (1987a, p.77,80) emended the diagnosis of this species. Size: length without operculum 40-60 µm, with opercula 14-16 µm longer, width 52-80 µm.