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Chatangiella madura

Chatangiella madura, Lentin and Williams, 1976, nom. subst. Chatangiella manumii, (Cookson and Eisenack, 1970), Lentin and Williams, 1976, non Chatangiella manumii, (Vozzhennikova, 1967), Lentin and Williams, 1976

Originally Deflandrea manumii, subsequently (and now) Chatangiella madura; see also Chatangiella manumii (combination illegitimate).

Holotype: Cookson and Eisenack, 1970, pl.11, fig.10
Locus typicus: Madura No. 1 Bore, Australia
Stratum typicum: Senonian

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Original description as Deflandrea manumi: [Cookson and Eisenack, 1970, p. 141-142]:

Description:
Shell relatively large, considerably longer than broad, divided approximately equally by a clearly-defined discontinuous girdle. The epitheca, which is slightly longer than the hypotheca, consists of a prominent apical region with strongly convex shoulder-like sides, a short, broad, median apical horn with a straight apex and outwardly slanting sides, and a slightly broader lower portion with convex sides which extends to the girdle. The sides of the hypotheca narrow gradually inwards towards a squarish antapex with a slightly pointed prominence on the right-hand side.
The girdle is relatively wide, strongly defined and broken at regular intervals.
The archeopyle is rather large and slightly angular in outline.
The wall of the shell is finely granular throughout. In addition, relatively large clearly-defined areas both above and below the individual subdivisions of the girdle, which resemble fields of Peridinium type (Manum 1963), are outlined by relatively prominent, closely arranged, bluntly-pointed solid thickenings which are circular in outline in surface view. The walls of the central body and shell are thin and in close contact.

Dimensions:
Holotype: c. 102 µm long, c. 60 µm broad.
Range: 20 specimens c. 95-138 µm long, c. 52-80 µm broad.

Affinities:
Both the shape and the size of the shell of D. manumi are closely similar to those of D. tripartita Cookson and Eisenack 1960, D. victoriensis Manum and Cookson 1964 and D. verrucosa Manum 1963, and show the Peridinium type of tabulation discussed by Manum in 1963. The fields outlined on the surface of the shell of D. manumi are 3", 4" and 5" and 2""", 3""" and 4""". Between the individual fields there are clearly marked longitudinal intercalary strips.

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Nohr-Hansen 1996, p. 32:

Remarks/Affinities:
The specimens from West Greenland often have very pronounced apical shoulders and a broad-based short apical horn. The periphragm is ornamented by grana or baculae, the ornament is most pronounced equatorially, but paratabulation is very seldom distinguished. The endocyst often has folded apical and antapical ends. The intercalary archeopyle is rounded iso-omegaform to ?iso-thetaform. The holotype of Chatangiella madura (Cookson & Eisenack, 1970) differs from the present material by having a more prominent ornamented periphragm reflecting the paratabulation and a round to spherical endocyst. Cookson & Eisenack (1970) mentioned that the shape and size of C. madura is close to that of C. tripartita, but that the paratabulation for the pre- and postcingular plate series on C. madura distinguishes the species from C. tripartita. Ionannides (1986) mentioned in his remarks on the C. madura specimens recorded from Bylot Island, arctic Canada, that C. madura and C. tripartita in gross morphology are very similar and they perhaps form part of the intraspecific variability of a single species.

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Occurrence:
In West Greenland Chatangiella sp. cf. C. madura has been recorded only from the Coniacian C. sp. cf. C. madura and Spinidinium echinoideum intervals from Svartenhuk Halv0. Cookson & Eisenack (1970) described C. madura from the Senonian of Western Australia. Ioannides (1986) recorded the species from the lower part of his Santonian-Campanian palynological interval I on Bylot Island, arctic Canada. Harker et al. (1990) recorded C. madura from a very narrow interval in the "latest" Campanian in the Western Interior of the U.S.A.. They recorded the almost identical species C. tripartita from the "latest" Santonian to "earliest" Maastrichtian (which included the majority of their studied stratigraphical interval). Kirsch (1991) reported C. madura from the lower part of his Coniacian/Santonian Raetiaedinium truncigerum Zone and from his Campanian Areoligera coronata Zone in Oberbayern, Germany. Nunez-Betelu (1994) recorded C. madura from his Zone 2 (late Turonian to late Coniacian) and his Zone 3 (late Coniacian to early Campanian) at Glacier Fiord, arctic Canada. Chatangiella tripartita was described by Cookson & Eisenack (1960) from probable late Turonian to middle Senonian of Australia. Aurisano (1989) recorded an early Santonian to early late Campanian range for C. tripartita in the New Jersey Atlantic Coastal Plains, U.S.A. Schi01er (1992) reported C. tripartita from the early to "mid"-Coniacian on the Danish island of Bornholm. Costa & Davey (1992) reported the C. tripartita - victoriensis complex from late Cenomanian to early Maastrichtian in the North Sea region. Marheinecke (1992) recorded the species as ranging up to the late part of the early Maastrichtian in Germany. Helby et al. (1987) reported C. tripartita from early Santonian (consistent) to middle and late Santonian (inconsistent).
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