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Chatangiella tripartita
Chatangiella tripartita, (Cookson and Eisenack, 1960), Lentin and Williams, 1976; Emendation as Deflandrea tripartita: Cookson and Manum, 1964, p. 521-522
Originally Deflandrea, subsequently Australiella, thirdly (and now) Chatangiella. This species is the type species of Australiella Vozzhennikova, 1967, which Lentin and Williams, 1976, considered to be a tax. jr. synonym of Chatangiella Vozzhennikova, 1967.
Holotype: Cookson and Eisenack, 1960, pl. 1, fig. 10; Cookson and Manum, 1964, pl.76, figs.1-2
Locus typicus: North of Gingin, Perth Basin, Western Australia
Stratum typicum: Senonian
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Original description as Deflandrea tripartiata: [Cookson and Eisenack, 1960, p. 2-3]:
Description:
Shell elongate, somewhat flat, divided approximately equally by a shallow, circular girdle, the borders of which form re-entrant angles on both sides. The middle portion of the shell, which is almost entirely filled with a large capsule, is widest at the girdle, narrowing from thence, rather abruptly, both proximally and distally. Above the upper limit of the capsule the shell again bulges prominently before terminating in a short, broad, bluntly pointed horn. The distal portion of the shell, i. e., the portion below the lower limit of the capsule, is straight sided, truncate, and prolonged on one side into a short pointed horn. The shell-membrane is rather sparsely, finely to coarsely granular. The capsular membrane is more finely and closely granular. The pylome is circular to hoof-shaped.
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Emended description as Deflandrea tripartiata:
Cookson and Manum, 1964, p. 521-522:
Description:
Emended on the basis of the holotype. Theca elongate, equally divided by an inconspicuous girdle. The middle portion of the theca, which is almost entirely filled with a large capsule, is widest in the region of the girdle and rather abruptly narrowed from thence towards both the upper and lower limits of the capsule. Above the upper limit of the capsule the theca bulges prominently before terminating in a short, broad, bluntly pointed horn. The theca-membrane is c. 1.0 µm thick and perfectly smooth in optical section; in surface view a fine pattern is faintly visible under an oil immersion lens (n.a. > 1.0 µm). The wall of the capsule, which is finely but distinctly granular, varies in thickness being c. 2.5 µm anteriorly and posteriorly and c. 1.0 µm where it is in contact with the theca. The intercalary archeopyle is somewhat elongate and trapezium-shaped.
Affinities:
The girdle in the holotype is ill-defined and mainly indicated by two pairs of short, low, parallel ridges, one on either side of the ventral surface at short distances from the lateral margins of the theca. The direction of these ridges indicates that the girdle was slightly laevo-rotatory. In the middle of the dorsal surface, a fold-like line which seems to represent a portion of the girdle and its borders is indicated. The apparent discontinuity of the girdle on the ventral surface suggests that it may have been "broken" in a manner similar to that characterizing certain other species of Deflandrea (Manum 1963), including D. victoriensis. As in these forms, the more prominent antapical horn in D. tripartita is on the left hand side of the hypotheca.
-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------Supplemental description:
Hultberg, 1985, p. 113-114:
Description:
Bicavate cyst, composed of endophragm and periphragm, separated by pericoels, but in contact at the paracingulum.
The surface of the endophragm is smooth. The surface of the periphragm is granulate. The shape of the endophragm is transversely ellipsoidal. The periphragm is elongate, with a prominent apical horn, and two, asymmetrically developed antapical horns. The left horn is always the largest. The epicyst bears prominent shoulders.
No trace of paratabulation is present, except the archeopyle.
The archeopyle is intercalary, type I, formed by the detachment of paraplate 2a. The archeopyle is narrow and slightly omegaform.
Paracingulum is indicated by folds in the periphragm. Also, because of the width of the endophragm, the periphragm bulges outwards in the paracingular region.
Parasulcus is not discernible.
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Notes:
G.L. Williams short notes on species, Mesozoic-Cenozoic dinocyst course, Urbino, Italy, May 17-22, 1999 - LPP VIEWER CD-ROM 99.5.
Chatangiella tripartita (Cookson and Eisenack, 1960a) Lentin and Williams, 1976, has a pericyst widest at the cingulum and well-developed shoulders. Antapically the pericyst sides in the area of the pericoel are straight., there is a short left antapical horn. The pericyst is finely to coarsely granular. The endocyst is more finely granular. The endocyst and pericyst are closely appressed laterally.
Size: pericyst 100-120 µm long, 59-71 µm wide. Endocyst 48-60 µm long, 48-60 µm wide.
Originally Deflandrea, subsequently Australiella, thirdly (and now) Chatangiella. This species is the type species of Australiella Vozzhennikova, 1967, which Lentin and Williams, 1976, considered to be a tax. jr. synonym of Chatangiella Vozzhennikova, 1967.
Holotype: Cookson and Eisenack, 1960, pl. 1, fig. 10; Cookson and Manum, 1964, pl.76, figs.1-2
Locus typicus: North of Gingin, Perth Basin, Western Australia
Stratum typicum: Senonian
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Original description as Deflandrea tripartiata: [Cookson and Eisenack, 1960, p. 2-3]:
Description:
Shell elongate, somewhat flat, divided approximately equally by a shallow, circular girdle, the borders of which form re-entrant angles on both sides. The middle portion of the shell, which is almost entirely filled with a large capsule, is widest at the girdle, narrowing from thence, rather abruptly, both proximally and distally. Above the upper limit of the capsule the shell again bulges prominently before terminating in a short, broad, bluntly pointed horn. The distal portion of the shell, i. e., the portion below the lower limit of the capsule, is straight sided, truncate, and prolonged on one side into a short pointed horn. The shell-membrane is rather sparsely, finely to coarsely granular. The capsular membrane is more finely and closely granular. The pylome is circular to hoof-shaped.
-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
Emended description as Deflandrea tripartiata:
Cookson and Manum, 1964, p. 521-522:
Description:
Emended on the basis of the holotype. Theca elongate, equally divided by an inconspicuous girdle. The middle portion of the theca, which is almost entirely filled with a large capsule, is widest in the region of the girdle and rather abruptly narrowed from thence towards both the upper and lower limits of the capsule. Above the upper limit of the capsule the theca bulges prominently before terminating in a short, broad, bluntly pointed horn. The theca-membrane is c. 1.0 µm thick and perfectly smooth in optical section; in surface view a fine pattern is faintly visible under an oil immersion lens (n.a. > 1.0 µm). The wall of the capsule, which is finely but distinctly granular, varies in thickness being c. 2.5 µm anteriorly and posteriorly and c. 1.0 µm where it is in contact with the theca. The intercalary archeopyle is somewhat elongate and trapezium-shaped.
Affinities:
The girdle in the holotype is ill-defined and mainly indicated by two pairs of short, low, parallel ridges, one on either side of the ventral surface at short distances from the lateral margins of the theca. The direction of these ridges indicates that the girdle was slightly laevo-rotatory. In the middle of the dorsal surface, a fold-like line which seems to represent a portion of the girdle and its borders is indicated. The apparent discontinuity of the girdle on the ventral surface suggests that it may have been "broken" in a manner similar to that characterizing certain other species of Deflandrea (Manum 1963), including D. victoriensis. As in these forms, the more prominent antapical horn in D. tripartita is on the left hand side of the hypotheca.
-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------Supplemental description:
Hultberg, 1985, p. 113-114:
Description:
Bicavate cyst, composed of endophragm and periphragm, separated by pericoels, but in contact at the paracingulum.
The surface of the endophragm is smooth. The surface of the periphragm is granulate. The shape of the endophragm is transversely ellipsoidal. The periphragm is elongate, with a prominent apical horn, and two, asymmetrically developed antapical horns. The left horn is always the largest. The epicyst bears prominent shoulders.
No trace of paratabulation is present, except the archeopyle.
The archeopyle is intercalary, type I, formed by the detachment of paraplate 2a. The archeopyle is narrow and slightly omegaform.
Paracingulum is indicated by folds in the periphragm. Also, because of the width of the endophragm, the periphragm bulges outwards in the paracingular region.
Parasulcus is not discernible.
-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
Notes:
G.L. Williams short notes on species, Mesozoic-Cenozoic dinocyst course, Urbino, Italy, May 17-22, 1999 - LPP VIEWER CD-ROM 99.5.
Chatangiella tripartita (Cookson and Eisenack, 1960a) Lentin and Williams, 1976, has a pericyst widest at the cingulum and well-developed shoulders. Antapically the pericyst sides in the area of the pericoel are straight., there is a short left antapical horn. The pericyst is finely to coarsely granular. The endocyst is more finely granular. The endocyst and pericyst are closely appressed laterally.
Size: pericyst 100-120 µm long, 59-71 µm wide. Endocyst 48-60 µm long, 48-60 µm wide.