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Cyclonephelium pastielsii
Cyclonephelium pastielsii Deflandre and Cookson, 1955
Now Glaphyrocysta. Originally Cyclonephelium, subsequently (and now) Glaphyrocysta.
The references to Pastiels, 1948, in Downie and Sarjeant, 1966, as pl.5, fig.6, and in Harker and Sarjeant, 1975, as p.47, pl.15, fig.5, are incorrect. The correct citation is Pastiels, 1948, pl.5, fig.15).
Holotype: Pastiels, 1948, pl.5, fig.15 (as Membranilarnacia cf. Iiradiscoides 0. Wetzel, 1933)
Locus typicus: Quenast, Blorquiau Quarry, Belgium
Stratum typicum: Early Eocene
Translation Pastiels, 1948, p. 47: LPP
Translation Gocht, 1969: Geological Survey of Canada
Description: Pastiels, 1948, p. 47: Membranilarnax cf. liradiscoides O. Wetzel
Central capsule lenticular, ornamented in the equatorial region with radiating appendices, simple or bifurcate, joined distally by a network of tubular ramifications forming largely areolate meshes. The reproduction of the plate description of M. cf. liradiscoides, given by O. Wetzel, 1933, pl.6, fig. 3-4, is too incomplete to attribute this specimen to that species.
Diameter of the capsule 50 Ám, total diameter approximately 80 Ám.
Eaton, 1976, p. 259: Cyclonephelium pastielsii
Dimensions: Holotype: cyst body (without operculum) 53x57 Ám; length of processes up to 30 Ám.
Remarks: As in the case of Cyclonephelium exuberans, Deflandre & Cookson
(1955: 285) also failed to designate a holotype for Cyclonephelium pastielsi, their new species name for forms described from the Lower Eocene of Belgium by Pastiels (1948) as Membranilarnax cf. Iiradiscoides 0. Wetzel 1933. However, since he only provided one illustration of M. cf. Iiradiscoides (Pastiels 1948: pl. 5, fig. 15, a probable camera lucida drawing) this must be the holotype, and it is here designated as such. Pastiels" original description of this form (1948: 47) indicates that it consists of a central body with processes restricted to the peripheral zone. The processes are simple or bifurcate, distally ramified and interconnected, and the illustrated specimen is devoid of them on reflected precingular plates 3"" and 6"".
In overall form the specimen illustrated by Pastiels is more closely comparable to Cyclonephelium retiintextum Cookson 1965 (De Coninck 1968: 41) than C. exuberans (Williams & Downie 1966: 225, 227). On this basis, C. pastielsi can only be considered to have been recorded from the Lower Eocene of Belgium (Pastiels 1948, as M. cf. Iiradiscoides; De Coninck 1965, 1968, Gruas-Cavagnetto 1968) and of northern France (Gruas-Cavagnetto 1968, 1970). There are also several listed records without illustration or description which cannot be commented on: Lower Eocene of Belgium (De Coninck 1967), of northern France (Chateauneuf & Gruas-Cavagnetto 1968) and of the Hampshire Basin in southern England (Gruas-Cavagnetto 1970); Middle Eocene of northern France (Gruas-Cavagnetto 1971).
Gocht, 1969, p.60,61:
The flattened central body has a 6-jagged apical archeopyle; apical calottes were not found. Processes are numerous, slender, standing separately or in fascicles. Distally they are joined into an irregular fine reticulum; the filaments of the reticulum sometimes expand into delicate, planar strips.
On the main body, the centers of the dorsal and ventral sides remain unoccupied. On the ventral side, the middle field plate (6"") does not bear any processes, but the lateral and dorsal precingulars bear groups of processes, which may join to form a peripheral cuff. They are usually directed forward. Below the unoccupied area, the processes of the postcingular series follow, of which at least the middle dorsal ones (2"""-4""") are distinctly grouped; they are directed posteriorly, toward the antapex. Distal connections between processes of the pre- and postcingular series are formed only laterally, not over the unoccupied central surfaces. On some specimens, also an antapical process group (1"""") is conspicuous. This structure conforms to the description and illustration of Williams and Downie, 1966; whereas the lectotype of the species (Pastiels, 1949) bears no processes also on the dorsal field 3"".
Occurrence: Early Eocene.
Now Glaphyrocysta. Originally Cyclonephelium, subsequently (and now) Glaphyrocysta.
The references to Pastiels, 1948, in Downie and Sarjeant, 1966, as pl.5, fig.6, and in Harker and Sarjeant, 1975, as p.47, pl.15, fig.5, are incorrect. The correct citation is Pastiels, 1948, pl.5, fig.15).
Holotype: Pastiels, 1948, pl.5, fig.15 (as Membranilarnacia cf. Iiradiscoides 0. Wetzel, 1933)
Locus typicus: Quenast, Blorquiau Quarry, Belgium
Stratum typicum: Early Eocene
Translation Pastiels, 1948, p. 47: LPP
Translation Gocht, 1969: Geological Survey of Canada
Description: Pastiels, 1948, p. 47: Membranilarnax cf. liradiscoides O. Wetzel
Central capsule lenticular, ornamented in the equatorial region with radiating appendices, simple or bifurcate, joined distally by a network of tubular ramifications forming largely areolate meshes. The reproduction of the plate description of M. cf. liradiscoides, given by O. Wetzel, 1933, pl.6, fig. 3-4, is too incomplete to attribute this specimen to that species.
Diameter of the capsule 50 Ám, total diameter approximately 80 Ám.
Eaton, 1976, p. 259: Cyclonephelium pastielsii
Dimensions: Holotype: cyst body (without operculum) 53x57 Ám; length of processes up to 30 Ám.
Remarks: As in the case of Cyclonephelium exuberans, Deflandre & Cookson
(1955: 285) also failed to designate a holotype for Cyclonephelium pastielsi, their new species name for forms described from the Lower Eocene of Belgium by Pastiels (1948) as Membranilarnax cf. Iiradiscoides 0. Wetzel 1933. However, since he only provided one illustration of M. cf. Iiradiscoides (Pastiels 1948: pl. 5, fig. 15, a probable camera lucida drawing) this must be the holotype, and it is here designated as such. Pastiels" original description of this form (1948: 47) indicates that it consists of a central body with processes restricted to the peripheral zone. The processes are simple or bifurcate, distally ramified and interconnected, and the illustrated specimen is devoid of them on reflected precingular plates 3"" and 6"".
In overall form the specimen illustrated by Pastiels is more closely comparable to Cyclonephelium retiintextum Cookson 1965 (De Coninck 1968: 41) than C. exuberans (Williams & Downie 1966: 225, 227). On this basis, C. pastielsi can only be considered to have been recorded from the Lower Eocene of Belgium (Pastiels 1948, as M. cf. Iiradiscoides; De Coninck 1965, 1968, Gruas-Cavagnetto 1968) and of northern France (Gruas-Cavagnetto 1968, 1970). There are also several listed records without illustration or description which cannot be commented on: Lower Eocene of Belgium (De Coninck 1967), of northern France (Chateauneuf & Gruas-Cavagnetto 1968) and of the Hampshire Basin in southern England (Gruas-Cavagnetto 1970); Middle Eocene of northern France (Gruas-Cavagnetto 1971).
Gocht, 1969, p.60,61:
The flattened central body has a 6-jagged apical archeopyle; apical calottes were not found. Processes are numerous, slender, standing separately or in fascicles. Distally they are joined into an irregular fine reticulum; the filaments of the reticulum sometimes expand into delicate, planar strips.
On the main body, the centers of the dorsal and ventral sides remain unoccupied. On the ventral side, the middle field plate (6"") does not bear any processes, but the lateral and dorsal precingulars bear groups of processes, which may join to form a peripheral cuff. They are usually directed forward. Below the unoccupied area, the processes of the postcingular series follow, of which at least the middle dorsal ones (2"""-4""") are distinctly grouped; they are directed posteriorly, toward the antapex. Distal connections between processes of the pre- and postcingular series are formed only laterally, not over the unoccupied central surfaces. On some specimens, also an antapical process group (1"""") is conspicuous. This structure conforms to the description and illustration of Williams and Downie, 1966; whereas the lectotype of the species (Pastiels, 1949) bears no processes also on the dorsal field 3"".
Occurrence: Early Eocene.