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Wilsonidium intermedium

Wilsonidinium intermedium (Cookson and Eisenack, 1961) Costa and Downie, 1979

Now Castellodinium. Originally Wetzeliella, subsequently Rhombodinium, thirdly Wilsonidium, fourthly (and now) Castellodinium.
Holotype: Cookson and Eisenack, 1961, pl.1, fig.5
Locus typicus: Rottnest Bore, W. Australia
Stratum typicum: Eocene

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G.L. Williams short notes on species, Mesozoic-Cenozoic dinocyst course, Urbino, Italy, May 17-22, 1999
Wilsonidium intermedium (Cookson and Eisenack, 1961b) Costa and Downie, 1979. According to Cookson and Eisenack (1961b, p.40), Wilsonidium intermedium has a somewhat longer than broad pericyst, rounded rhombohedric in outline, and a broadly rounded apex with one or two spines. The lateral horns are represented by slight, broadly rounded or indented expansions of the pericyst. There is a single, short, pointed antapical horn. Cingulum indicated by low ledges or folds, especially at the lateral margins, or by two rows of short teeth. The margins of the pericyst are smooth, or slightly wavy or toothed. The dorsal and ventral surfaces are smooth or sparingly dotted, the dots sometimes outlining fields. Endocyst large, filling most of pericyst but with ambital pericoel. Wilsonidium intermedium is intermediate between Wilsonidium lineidentatum and Rhombodinium glabrum. Size: length 111-143 µm, width 97-130 µm, internal body of type 100 x 100 µm.
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Original diagnosis: Cookson and Eisenack, 1961, p.40: Wetzeliella (Rhombodinium) intermedia
Shell flat, usually somewhat longer than broad, rounded rhombic in outline with slightly convex to nearly straight sides and a broadly rounded apex with one or two centrally placed teeth. The lateral horns are represented by slight, broadly-rounded or indented expansions of the outer membrane; a single short pointed horn is developed to one side of the antapex. A circular girdle which crosses the shell in its broadest region is indicated by low ledges or folds especially at the lateral margins or by two rows of short teeth. The margins of the shell are smooth or slightly wavy or toothed, the dorsal and ventral surfaces are either smooth or sparingly dotted, the dots sometimes outliningfields comparable with those of W. lineidentata. The internal body is large, flat, and oval in outlines its wall is thin and inoptical section is seento be composedof smallclosely arranged rods. A large squarish pylome is situated in the epitheca.
Dimensions: Type - length 125 µm, breadth 122 µm, internal body 100 x 100 µm. Range length 111 - 143 µm, breadth 97 - 130 µm.

Affinities:
Cookson and Eisenack, 1961, p.40: Wetzeliella (Rhombodinium) intermedia Wetzeliella intermedia as its name suggests, seems to be a form intermediate between W. lineidentata and W. glabra Cookson. The slight ly denticulate margins and surface thickenings, sometimes linearly arranged so as to enclose fields, indicate a certain relationship with W. lineidentata, on the other hand the practically smooth margin and surface of other examples suggests an affinity with W. glabra.
W. intermedia agrees with W. rhomboidea Alberti (1961) in certain features, but the latter species has no indication of spines.
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