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Pyxidiella simplex

Pyxidiella? simplex Harland, 1979

Originally (and now) Pyxidiella?, subsequently Tectatodinium. Harland, 1979 questionably included this species in Pyxidiella. Edwards, 1984 transferred this species to Tectatodinium Wall, 1967; however, Lentin and Williams, 1985 questionably retained it in Pyxidiella.

Holotype: Harland 1979b: Plate 3, Fig. 12
Locus typicus: Bay of Biscay, DSDP Leg 48 (Hole 400A), eastern North Atlantic
Stratum typicum: late Miocene
Age: Late Miocene

Original diagnosis: Harland 1979b, p. 537-538
Ovoid to rounded pentagonal cyst made up of two wall layers closely adpressed, and carrying a microgranulate, granulate, conate, or spinate to foveolate ornamentation. Wall thick (2.0µm) and may be drawn out into a rounded apical boss. Tabulation not present, archeopyle probably intercalary presumably formed by the loss of plate 2a.

Dimensions: Holotype: Length 36.0 µm; breadth 30.0 µm; Range: Length 32.0-46.0 µm; breadth 28.0-38.0 µm. Four specimens measured.

Original description: Harland 1979b, p. 537-538
Cyst shape is variable, some specimens showing a clearly pentagonal outline (Plate 3, Figure 12) whilst others are ovoid (Plate 3, Figures 13-15). The ornamentation is often difficult to decipher, but probably both negative and positive elements are present; some specimens carry distinct cones or granules. The archeopyle shape is of interest as it suggests the quadra-type peridinacean plan, this together with position on. the cyst indicate an archeopyle of the intercalary type (Plate 3, Figure 12).

Remarks: This cyst is most easily attributed to the genus Pyxidiella Cookson and Eisenack and is similar to P. scrobiculata (Deflandre and Cookson) Cookson and Eisenack, 1958, from which it differs in not possessing such a coarse positive ornament, in being half the size and in sometimes possessing a pentagonal outline.

Occurrence: Observed in late Miocene sediments from Hole 400A.
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