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Spiniferites adnatus

Spiniferites adnatus Matsuoka and Bujak, 1988

Holotype: Matsuoka and Bujak, 1988, pl.10, fig.8a-b
Paratype: Matsuoka and Bujak, 1988
Locus typicus: Norton Sound COST No. 1 Well, Norton Sound, Bering Sea
Stratum typicum: Late Oligocene

Original diagnosis: Matsuoka and Bujak, 1988, p.73
Intermediate ovoidal to elongate proximochorate cyst. Epicyst hemi-ellipsoidal and always longer than hypocyst, with or without apical boss. Cyst wall thin; periphragm smooth to chagrinate, endophragm smooth. Processes mostly gonal, membranous or sometimes hollow, short cylindrical to tubiform with irregularly foliate, furcate or rarely secate distal extremities. Parasutural septa distinctive, low, sometimes membranous and representing the paratabulation 4", 6"", 6c, 6""", 1p, 1"""" and xs. Parasutural septa between paraplates 1"""", 3"""-4""", and between paraplates 1" and 2" often membranous with branched to entire distal ends. Paracingulum clear, almost circular and comprising six equatoriallyexpanded rectangular paraplates; parasulcus delimited by parasutural septa and almost straight. Archeopyle simple precingular formed by the loss of paraplate 3".

Original description: Matsuoka and Bujak, 1988, p.73-74
The cyst body varies from ovoidal to ellipsoidal and the epicyst is longer than the hypocyst. The cyst wall comprises a periphragm and endophragm adpressed between the processes. The processes are mostly gonal, membranous, cylindrical to tubiform, with extremities that vary from bifurcate to irregularly foliate. Some of the gonal processes are hollow and distally secate. The parasutures are distinctive and often membranous between the 1"""" and 3"""-4""" paraplates, and sometimes between the 1" and 2" papaplates. These membranous parasutural septa arehystricate to entire. The paracingulum is approximately circular and the parasulcus is almost straight. The archeopyle is precingular, reduced, and the operculum is monoplacoid (3" ) and detached.
Dimension: Holotype; length of cyst 53 µm, width 45 µm, length of processes up to 9 µm, Range; length of cyst 39-53 µm, width 33-46 µm, length of processes 8-12 µm. Number of specimens measured; 10.
Thecal affinities: Unknown; possibly related to the genus Gonyaulax.

Matsuoka and Bujak, 1988: Spiniferites adnatus resembles S. nortonensis sp. nov. in possessing membranous gonal processes and parasutural septa, but differs in having a more elongate cyst body and longer processes. S. adnatus is also similar to S. ellipsoideus, S. membranaceus (Rossignol) and S. mirabilis (Rossignol), but is distinguished from S. ellipsoideus in having a larger cyst body and hollow gonal processes, from S. membranaceous in possessing an ovoidal cyst body and hollow processes with foliate distal extremities, and from S. mirabilis in possessing only gonal and sometimes hollow processes.
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