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Spiniferites incertus

Spiniferites incertus (Klumpp, 1953, p.389–390, pl.17, figs.1–2) Sarjeant, 1981, p.109–110. Emendations: Morgenroth, 1966a, p.15, as Hystrichosphaera incerta; Sarjeant, 1981, p.109–110, as Spiniferites incertus.

Originally Areoligera, subsequently Hystrichosphaera, thirdly (and now) Spiniferites.
Tax. jr. synonym of Hystrichosphaera (now Spiniferites) pseudofurcata Klumpp, 1953, according to Gocht, 1969. Sarjeant, 1981, retained Hystrichosphaera (as Spiniferites) incerta as a separate species.
Morgenroth, 1966, emended the diagnosis of this species, as Hystrichosphaera incerta.

Holotype: Klumpp, 1953, pl.17, figs.1-2
Age: Late Eocene
Translation Klummp, 1953: Sarjeant, 1981, p. 109


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Original Diagnosis: Klummp, 1953, p. 389: Areoligera incerta
A species of the genus Areoligera with the following distinguishing characteristics: all plates irregularly quadrate without a distinct zone with elongate plates. The edge of the flat to concave fields is set with perpendicular spines, not with a velum.

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Emended diagnosis: Morgenroth, 1966a:

Nomenclature correction: The genus Hystrichokibotium KLUMPP 1953 includes Hystrichospheres with a feldsen capsule, which are said to be distinguished by the absence of a girdle region. However, the original type species, Hystrichokibotium pseudofurcatum KLUMPP, is a Hystrichosphaera whose girdle region appears at the edge due to its unfavorable location and could therefore be overlooked. In the paratypoids, the girdle is sometimes clearly identifiable, as is the case in the Lower Eocene material from Katharinenhof. This Hystrichosphaera species was described in detail by EISENACK & GOCHT 1960 as Hystrichosphaera tertiaria, although no specialisms can be found compared to the specimens described by KLUMPP under Areoligera incerta, Hystrichokibotium pseudofurcatum, and Hystrichosphaera furcata. This results in the situation where the genus name (Areoligera) is omitted from KLUMPP's version, and the species name (tertiaria) from EISENACK & GOCHT's version for priority reasons.
Note: Whether the species "Hyslrichokibotium trabeculiferum" (DEFLANDRE & COOKSON 1955) is distinguished by the absence of a girdle line requires further investigation. The specimen illustrated by DEFLANDRE & COOKSON 1955 on Plate 8, Fig. 6, very closely resembles several abnormal specimens of Hystrichosphaera furcata (EHRENB.) found in the Lower Eocene of Fehmarn (cf. Plate 7, Fig. 5).
Dimensions: Capsule Ø 55μ (48μ/64μ), largest pylome Ø 20μ-28μ, specimen Ø 100μ-125μ. Process length 25μ–33μ (50 measurements). Number of specimens examined: 1,700.

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Emended diagnosis: Sarjeant, 1981, p. 109-110
Cyst broadly ovoidal to rounded-subpolygonal, chorate, spiniferate. Processes gonal in situation, relatively short (approximately one-third of the short diameter in length). Their form is highly irregular; they may consist of single, irregularly branched and fenestrate processes, single ramifying processes, or tight groups of processes, proximally close-set or sharing a common base, bifurcating at or near mid-point or linked by short cross-connections, distally capitate, symmetrically or asymmetrically bifid or (rarely) foliate. Low sutural crests, sometimes continuos and solid, sometimes branching and anastomosing, outline the following paratabulation: ?4", 0a, 6"", ?5c, 6""", 0-?1p, 1"""". Cingulum unusually broad. Archaeopyle precingular (type P), formed by loss of paraplate 3""; operculum normally free.
Dimensions: Holotype Iength 70 µm, breadth 60 µm, length of processes c.18-24 µm.
Description: The processes are extremely irregular in form and are indeed reminiscent of the component elements of the process clusters of Systematophora and allied genera. However, they arise only at paraplate junctions; sutural processes are absent. The surface of the phragma is finely and irregularly punctate.
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