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Phoberocysta neocomica ssp. neocomica
Phoberocysta neocomica ssp. neocomica (Gocht, 1957) Millioud, 1969; emend. Helby,1987
Autonym. Now redundant.
Taxonomic junior synonyms:
Wetzeliella? neocomica forma circulata (subsequently Phoberocysta neocomica subsp. circulata),
Wetzeliella? neocomica forma cruciformis (subsequently Phoberocysta neocomica subsp. cruciformis) and
Wetzeliella? neocomica forma dedecosa (subsequently Phoberocysta neocomica subsp. dedecosa), by implication in Monteil (1991b, p.477), who listed these taxa as taxonomic junior synonyms of Phoberocysta neocomica.
Holotype: Gocht, 1957, pl.19, fig.1; text-fig.7
Locus typicus: Ruehlertwist, Emsland, NW Germany
Stratum typicum: Hauterivian, Early Cretaceous
Translation Gocht, 1957: LPP
Original description: Gocht 1957, p. 172-176: Wetzeliella ?neocomica.
Diagnosis: Shell flattened, extremely variable in shape, generally more or less 4-5 sided, delicate. Closed edge rare; Shell all around mostly developing into bilayered and exteriorly closed projections, which may be located at several levels, but which are essentially limited to the region of the outer margin. Among them, an apical horn, two lateral horns, and an antapical horn smay often be clearly distinguished. Projections and horns often provided with pointed, blunt or trumpet shaped, flat protruberances or processes, which almost exclusively stand along the outline, and rarely on the surface. On the front and back sides projections or processes are sporadic. Along the margins of the apical and antapical horns, the protruberancesare are often arranged alternatively. Lateral horns mostly slightly squat, broad-based, sometimes slightly bent, giving rise to a slightly V-shaped cross-section.
Projections and processes may be considerably reduced or extremely enlarged, as well as the horns, which may also partly be missing or be unrecognizable as such.
Exceptionally, denticulate ridges or idications of a transverse furrow occur on the shell.
Inner body delicate, with a more or less oval outline.
Emended description: Helby, 1987, p. 310-313
Proximochorate cyst, outline rhomboidal with a prominent apical horn, cingular horns, left antapical horn and a periphragm bulge of variable size constituting a right antapical horn. Cingular horns variably developed, usually expressed as stout lateral bosses. Each cingular horn consisting of parts of 2 modified paraplates from each of the precingular and postcingular series and a cingular paraplate (2c) on the left and 2 cingular paraplates (4c and 5c) on the right. Posterior surface of the lateral tip of the horns, comprising the lateral extension of the postcingular paraplates, often inclined posteriorly. Lateral edges of the cingular horns slope anteriorly towards the cyst, although parasutural ridges give it a distinctive notched appearance, often exaggerated by development of substantial gonal spines. Left antapical horn prominent, usually rounded at the distal end. Cyst 2-layered, cornucavate, but wall layers closely appressed over most of the dorsal and ventral surfaces. Endocyst rhomboidal with small protrusions at horn bases apart from the right antapical horn. Periphragm varying from smooth to finely granulate, scabrate and occasionally perforate. Periphragm bearing a variety of spines and spinules. Large, solid, capitate spines (tapered from a very broad base, and up to 15µ long), generally with small spinules.
Spines generally gonal and parasutural and only occasionally intratabular. Spinules (0.5-1µ high) concentrated at the distal extremities of large spines and horns (particularly the apical and antapical horns) or may occur as scattered intratabular groups, or may be aligned parasuturally. Endophragm usually granulate. Paratabulation indicated by archeopyle, by the location of gonal and intergonal spines and occasionally by less distinct parasutural features such as delicate differentiation of the periphragm (scabration). Paratabulation formula interpreted as 4", 0a, 6"", 6c(?), 6""", 1p, 1"""" (fig.15). Archeopyle apical, type [tA], principle archeopyle suture zigzag with parasulcal notch set off to left side. Paracingulum transverse feature across dorsal surface onto paracingular horns where it is evident as a distinct concavity (exaggerated by gonal spine development) at the lateral edge of the horns. Parasulcus sinous, offset from a midventral position on the hypocyst to a left ventral position on the epicyst. Parasulcal plates not well defined.
Autonym. Now redundant.
Taxonomic junior synonyms:
Wetzeliella? neocomica forma circulata (subsequently Phoberocysta neocomica subsp. circulata),
Wetzeliella? neocomica forma cruciformis (subsequently Phoberocysta neocomica subsp. cruciformis) and
Wetzeliella? neocomica forma dedecosa (subsequently Phoberocysta neocomica subsp. dedecosa), by implication in Monteil (1991b, p.477), who listed these taxa as taxonomic junior synonyms of Phoberocysta neocomica.
Holotype: Gocht, 1957, pl.19, fig.1; text-fig.7
Locus typicus: Ruehlertwist, Emsland, NW Germany
Stratum typicum: Hauterivian, Early Cretaceous
Translation Gocht, 1957: LPP
Original description: Gocht 1957, p. 172-176: Wetzeliella ?neocomica.
Diagnosis: Shell flattened, extremely variable in shape, generally more or less 4-5 sided, delicate. Closed edge rare; Shell all around mostly developing into bilayered and exteriorly closed projections, which may be located at several levels, but which are essentially limited to the region of the outer margin. Among them, an apical horn, two lateral horns, and an antapical horn smay often be clearly distinguished. Projections and horns often provided with pointed, blunt or trumpet shaped, flat protruberances or processes, which almost exclusively stand along the outline, and rarely on the surface. On the front and back sides projections or processes are sporadic. Along the margins of the apical and antapical horns, the protruberancesare are often arranged alternatively. Lateral horns mostly slightly squat, broad-based, sometimes slightly bent, giving rise to a slightly V-shaped cross-section.
Projections and processes may be considerably reduced or extremely enlarged, as well as the horns, which may also partly be missing or be unrecognizable as such.
Exceptionally, denticulate ridges or idications of a transverse furrow occur on the shell.
Inner body delicate, with a more or less oval outline.
Emended description: Helby, 1987, p. 310-313
Proximochorate cyst, outline rhomboidal with a prominent apical horn, cingular horns, left antapical horn and a periphragm bulge of variable size constituting a right antapical horn. Cingular horns variably developed, usually expressed as stout lateral bosses. Each cingular horn consisting of parts of 2 modified paraplates from each of the precingular and postcingular series and a cingular paraplate (2c) on the left and 2 cingular paraplates (4c and 5c) on the right. Posterior surface of the lateral tip of the horns, comprising the lateral extension of the postcingular paraplates, often inclined posteriorly. Lateral edges of the cingular horns slope anteriorly towards the cyst, although parasutural ridges give it a distinctive notched appearance, often exaggerated by development of substantial gonal spines. Left antapical horn prominent, usually rounded at the distal end. Cyst 2-layered, cornucavate, but wall layers closely appressed over most of the dorsal and ventral surfaces. Endocyst rhomboidal with small protrusions at horn bases apart from the right antapical horn. Periphragm varying from smooth to finely granulate, scabrate and occasionally perforate. Periphragm bearing a variety of spines and spinules. Large, solid, capitate spines (tapered from a very broad base, and up to 15µ long), generally with small spinules.
Spines generally gonal and parasutural and only occasionally intratabular. Spinules (0.5-1µ high) concentrated at the distal extremities of large spines and horns (particularly the apical and antapical horns) or may occur as scattered intratabular groups, or may be aligned parasuturally. Endophragm usually granulate. Paratabulation indicated by archeopyle, by the location of gonal and intergonal spines and occasionally by less distinct parasutural features such as delicate differentiation of the periphragm (scabration). Paratabulation formula interpreted as 4", 0a, 6"", 6c(?), 6""", 1p, 1"""" (fig.15). Archeopyle apical, type [tA], principle archeopyle suture zigzag with parasulcal notch set off to left side. Paracingulum transverse feature across dorsal surface onto paracingular horns where it is evident as a distinct concavity (exaggerated by gonal spine development) at the lateral edge of the horns. Parasulcus sinous, offset from a midventral position on the hypocyst to a left ventral position on the epicyst. Parasulcal plates not well defined.