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Epiplosphaera reticulospinosa
Epiplosphaera reticulospinosa Klement, 1960, p.75-76
Taxonomic junior synonym: Sentusidinium (subsequently Epiplosphaera) ornatum, according to Courtinat (1989, p.176) — however, Poulsen (1992a, p.68) considered Epiplosphaera ornata to be a taxonomic junior synonym of Epiplosphaera gochtii.
Holotype: Klement, 1960, pl.8, figs.10–11; Fauconnier and Masure, 2004, pl.27, figs.6–8.
Paratypes: Klement, 1960
Locus typicus: Well Scherstetten 1, SW Germany
Stratum typicum: Malm Gamma 3 (Similis-beds), Early Kimmeridgian
Translation Klement, 1960: LPP, GSC
Original diagnosis: Klement, 1960, p. 75-76
A species with the higher-grade features of the genus Epiplosphaera, characterized by a squat-ellipsoidal shell with a clearly expressed, irrularly polygonal, fine-meshed basal network, developed by low ridges. Processes simple thorn-like, rising from junctions of the network, standing mostly isolated on upper and lower side, marginally however connected by low, rigid ridges, which show a meridian course.
Dimensions: Holotype: shell with processes: L:B 69:69 Ám, shell without processes L:B 52:49 Ám, process length 12 Ám.. Paratypes: shell with processes: L:B 63:62 and 69:60 Ám, shell without processes L:B 55:49 and 52:46 Ám, process length 12 and 15 Ám.
Original description: Klement 1960, p. 75: Epiplosphaera reticulospinosa
This species likewise was not found very frequently, but was preserved in an excellent manner, permitting a subtle observation of the finest details.
The very prominent, small-meshed, irregular polygonal basal network (with mesh-widths ranging from 4 to 7 microns) bears relatively short, spiny processes (with broadened basal regions) in some of its junction points. On the upper-, and bootom-side of the shell, said processes usually stand isolated, whereas marginally, they are connected with one anotherby means of stiff sutures, which extend in meridian direction and hang in a concave manner between the processes. Furthermore, when examining the specimens, one gains the impression as if the marginalmeshes of the basic network were approxiamtley of the same size and were lying parallel to one another.
In all representatives of the species here under discussion, one pole of the shell appeared to be split-off in each case in a similar calotte-like manner. This characteristic feature is considered to be related to the pylome formation.
The color of the thin-walled shell membrane, and that of the sutures, is a light yellow.
Relationships: Klement 1960, p. 75
The representatives of the present species can be differentiated very well from the E. bireticulata by the absence of the closed sutural network structures on the shell surface, on the one hand, and by the simple, spine-like processes and by the stiffness of their interconnecting sutures. As far as the E.areolata is concerned, the precise delimitation from the present species is based on the fact that, in the case of the E. areolata, the interconnecting structures between the individual processes never are present in form of sutures, but solely in form of bulges and that said interconnecting structures moreover join together to form closed formations.
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Poulsen, 1992, p. 68
Remarks: The length of the spines and the height of the crests ranges from 15-12 Ám in the type material to 9 Ám in the Polish material.
Stratigraphy:
Poulsen, 1992, p. 68, 70
Early Oxfordian to the earliest Volgian/Tithonian. In the studied material, E. reticulospinosa occurs from the Middle Oxfordian (Transversarium Zone) to the Late Kimmeridgian (Eudoxus-Autissiodorensis Zones). The species has been recorded from the Earlv Oxfordian to the Late Kimmeridgian (Eudoxus Zone) of Germany (Brenner, 1988; D³rr, 1988; Klement, 1960). It has been recorded from the Middle Oxfordian? to the earliest Tithonian of France (Courtinat, 1989; loanides et al., 1988). It is recorded in the Late Oxfordian to mid Kimmeridgian (Serratum to Mutabilis Zones) of England (N÷hr-Hansen, 1986; Thomas and Cox, 1988).
Taxonomic junior synonym: Sentusidinium (subsequently Epiplosphaera) ornatum, according to Courtinat (1989, p.176) — however, Poulsen (1992a, p.68) considered Epiplosphaera ornata to be a taxonomic junior synonym of Epiplosphaera gochtii.
Holotype: Klement, 1960, pl.8, figs.10–11; Fauconnier and Masure, 2004, pl.27, figs.6–8.
Paratypes: Klement, 1960
Locus typicus: Well Scherstetten 1, SW Germany
Stratum typicum: Malm Gamma 3 (Similis-beds), Early Kimmeridgian
Translation Klement, 1960: LPP, GSC
Original diagnosis: Klement, 1960, p. 75-76
A species with the higher-grade features of the genus Epiplosphaera, characterized by a squat-ellipsoidal shell with a clearly expressed, irrularly polygonal, fine-meshed basal network, developed by low ridges. Processes simple thorn-like, rising from junctions of the network, standing mostly isolated on upper and lower side, marginally however connected by low, rigid ridges, which show a meridian course.
Dimensions: Holotype: shell with processes: L:B 69:69 Ám, shell without processes L:B 52:49 Ám, process length 12 Ám.. Paratypes: shell with processes: L:B 63:62 and 69:60 Ám, shell without processes L:B 55:49 and 52:46 Ám, process length 12 and 15 Ám.
Original description: Klement 1960, p. 75: Epiplosphaera reticulospinosa
This species likewise was not found very frequently, but was preserved in an excellent manner, permitting a subtle observation of the finest details.
The very prominent, small-meshed, irregular polygonal basal network (with mesh-widths ranging from 4 to 7 microns) bears relatively short, spiny processes (with broadened basal regions) in some of its junction points. On the upper-, and bootom-side of the shell, said processes usually stand isolated, whereas marginally, they are connected with one anotherby means of stiff sutures, which extend in meridian direction and hang in a concave manner between the processes. Furthermore, when examining the specimens, one gains the impression as if the marginalmeshes of the basic network were approxiamtley of the same size and were lying parallel to one another.
In all representatives of the species here under discussion, one pole of the shell appeared to be split-off in each case in a similar calotte-like manner. This characteristic feature is considered to be related to the pylome formation.
The color of the thin-walled shell membrane, and that of the sutures, is a light yellow.
Relationships: Klement 1960, p. 75
The representatives of the present species can be differentiated very well from the E. bireticulata by the absence of the closed sutural network structures on the shell surface, on the one hand, and by the simple, spine-like processes and by the stiffness of their interconnecting sutures. As far as the E.areolata is concerned, the precise delimitation from the present species is based on the fact that, in the case of the E. areolata, the interconnecting structures between the individual processes never are present in form of sutures, but solely in form of bulges and that said interconnecting structures moreover join together to form closed formations.
----------------------------------------------------------
Poulsen, 1992, p. 68
Remarks: The length of the spines and the height of the crests ranges from 15-12 Ám in the type material to 9 Ám in the Polish material.
Stratigraphy:
Poulsen, 1992, p. 68, 70
Early Oxfordian to the earliest Volgian/Tithonian. In the studied material, E. reticulospinosa occurs from the Middle Oxfordian (Transversarium Zone) to the Late Kimmeridgian (Eudoxus-Autissiodorensis Zones). The species has been recorded from the Earlv Oxfordian to the Late Kimmeridgian (Eudoxus Zone) of Germany (Brenner, 1988; D³rr, 1988; Klement, 1960). It has been recorded from the Middle Oxfordian? to the earliest Tithonian of France (Courtinat, 1989; loanides et al., 1988). It is recorded in the Late Oxfordian to mid Kimmeridgian (Serratum to Mutabilis Zones) of England (N÷hr-Hansen, 1986; Thomas and Cox, 1988).