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Gochteodinia villosa

Gochteodinia *villosa (Vozzhennikova, 1967, p.56, pl.12, figs.1a-b,2a-b,3a-b; pl.13, figs.1a-e,2,3a-d; pl.14, figs.1a-e,2a-i; pl.15, figs.1-2) Norris, 1978, p.7. Emendations: Lentin and Vozzhennikova, 1990, p.85 and Below, 1990, p.50, both as Gochteodinia villosa.

Originally Imbatodinium villosum, subsequently Pareodinia villosa (Vozzhennikova) (combination not validly published), thirdly Pareodinia dasyforma, fourthly (and now) Gochteodinia villosa.

Holotype: Vozzhennikova, 1967, pl.12, figs.3a-b; pl.15, fig.1; Eisenack and Kjellström, 1975b, figure to right - p.726f; Lentin and Vozzhennikova, 1990, pl.11, figs.7-8; text-fig.47; Fensome et al., 1995, fig.1 - p.1903; Fensome et al., 1996, figs.1-3 - p.2431.
Locus typicus: Moscow region, Russia
Stratum typicum: Late Jurassic
Translation Vozzhennikova, 1967: Lees in Sarjeant 1971 (Ed.)
Translation Below, 1990: LPP

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G.L. Williams short notes on species, Mesozoic-Cenozoic dinocyst course, Urbino, Italy, May 17-22, 1999 - LPP VIEWER CD-ROM 99.5.

Gochteodinia villosa (Vozzhennikova, 1967) Norris, 1978, emend. Lentin and Vozzhennikova, 1989, emend. Below, 1990. Below (1990, p.50) emended the diagnosis. The tabulation is as for the genus. The cyst is proximate, acavate-cornucavate, cylindrical, with the apex drawn out into an apical horn. Surface with numerous solid, cylindrical, distally cauliflorate, capitate, corynate, or evexate processes, having a maximum length equal to two-thirds of the diameter of the central body, but which can also be reduced to short, cauliflorate, capitate, evexate, cylindrical stumps. Archeopyle formed from loss of two anterior intercalary paraplates. Operculum solvate, opercular pieces secate. Size: length 71-100 µm, width 27-41 µm, length of apical horn 8 µm, process length 10-14 µm.
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Original description: Vozzhenikova 1967, p. 56: Imbatodinium villosum
Diagnosis: Theca pyriform, with convex sides. Transverse furrow deep. Longitudinal furrow S-shaped, extending from the base of the apical process to the antapex. Epitheca 2.5 to 3 times as big as the hyptoheca. Epitheca with an apical finger-like process bearing a slender cilium, and covered with short slender hairs (Pl. 12, fig. 1b, 2b, 3a). Hypotheca with unequal antapical processes, one of which
is broadly rounded, the other smaller and more bluntly rounded is sometimes beak-like (Pl. 13, fig. 3b, c, d). Theca covered with slender hairs which bifurcate at the distal end.
Dimensions: length: 70.5-100 Ám; breadth: 27-40.5 Ám, length of apical process 10.5-13.5 Ám.

Supplemental description: Norris 1978, p. 7: Gochteodinia villosa
Proximo-chorate cysts with a well-developed apical horn and elongated main body, both ornamented with non-tabular acuminate or bluntly terminated processes, which in length are approximately one quarter the main body equatorial diameter. Archeopyle 2I or 3I.

Emended description: Lentin and Vozzhennikova, 1990, p.85
Cyst shape elongate oval with one apical horn, without antapical horns, antapex rounded. Autophragm is ornamented with long slender, acuminate, bifurcate or clubshaped processes which are randomly distributed over the surface of the cyst; processes increase in length towards the antapex. The anterior tip of the apical horn is ornamented with a single, long, solid process. Paratabulation indicated only by location of the archeopyle. Archeopyle 2I, resulting from the loss of two intercalary paraplates, operculum free. Paracingulum and parasulcus not indicated.
Size: Holotype length 108 Ám, width 30 Ám, length of processes 10-14 Ám. Range, length 70.5-108 Ám width 27-41 Ám.

Emended decription: Below, 1990, p. 50
Vesicle/plate arrangement on amphiesma cop, pop, cap, 3', 2a, 6'', 7c, 6''', 2'''', as, FM, ls, rs, ps; 1' exsert, 1a and 2a pentagonal. Growth of thecal plates ?gonyaulacoidal.
Cyst proximate, acavate- cornucavate, cylindrical, apically evenly streched into apical horn, antapex rounded, equatorial section circular; wall consisting of thin pedium and densely prismatic-granular luxuria; surface smooth and disorderly covered with intraareate, peniareate and finate, solid, cylindrical, distally cauliflorate, capitate, corynate or evexate processes, varying in length, up to 2/3 of the central body-diameter,that may be reduced to short, cauliflorate, capitate, evexate, cylindrical stumps, nonareate except for the archaeopyle sutures; Paratabulation XPR, NR, 1a(arch)-2a(arch), NR'', NRc, NR''', NR'''', NRs; penta 1a (DDL) and 2a (DR), anteriorly geniculate 2'' (L), 4'' (DDR) and 5'' (R), anteriorly linear 3'' (DDL); archaeopyle anterior intercalary, 1a+2a; operculum solvate, opercular plates secate, general opercular formula 1a(s)+2a(s).
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