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Muderongia testudinaria

Muderongia testudinaria Burger, 1980; emend. Monteil, 1991

Originally (and now) Muderongia, subsequently Pseudomuderongia.
Tax. sr. synonym of Phoberocysta lowryi Backhouse, 1987, according to Monteil, 1991.
Tax. sr. synonym of Phoberocysta burgeri Helby, 1987, according to Monteil, 1991.
Tax. sr. synonym of Phoberocysta edgellii Helby, 1987, according to Monteil, 1991.
Jain and Khowaja-Ateequzzaman, 1984, transferred this species to Pseudomuderongia Jain and Khowaja-Ateequzzaman, 1984. Helby (1987, p.305) and Stover and Williams, 1987, retained it in Muderongia.

Holotype: Burger, 1980, fig.9B
Locus typicus: Carpentaria Basin, Australia
Stratum typicum: ?Hauterivian

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G.L. Williams short notes on species, Mesozoic-Cenozoic dinocyst course, Urbino, Italy, May 17-22, 1999

Muderongia testudinaria Burger, 1981b, emend. Monteil, 1991b. Monteil (1991b, p.476) provided the following emendation. Cysts proximate to proximochorate, compressed dorsoventrally, two-layered, cornucavate to delphicavate or circumcavate. Pericyst ceratioid, symmetrical, with 5 prominent tapered horns: 1 apical (axial, type AP I); 2 subequal lateral (bent, type L II), indented; 2 subequal antapical (not joined, type ATP III). Sometimes a “true” elbow may be developed on the postcingular extension... Distal extremities of horns closed and pointed; proximal extremities narrow to wide. Endocyst oval to rhombic, or pentangular, often with protrusions at the base of the horns. Periphragm thin, psilate to scabrate, sometimes locally finely perforated and/or folded; periphragm may be ornamented by processes (stages I to IV). Endophragm thin, psilate to granulate. Archeopyle apical, type (4A), with angular margin. Parasulcal notch marked. Operculum free. Paratabulation indicated by archeopyle, by low parasutural ridges or septa, and more or less clearly expressed by distribution and type of processes (stages I to IV), when those are present. Occasionally, periphragm showing folds or penitabular microperforations. Formula: ?pr, 4', 0a, 6", 6c, 6"’, 1p, 1"”, ?xs. Pericingulum marked by indentation of lateral horns and by parallel transverse structures. Perisulcus sinuous, marked by parasulcal notch offset. Size: total length (two specimens) 150, 154 µm, length of 11 dehisced specimens 80-110 µm, endocyst 59-77 µm long (dehisced), and 57-96 µm wide, antapical horns 20-38 µm long, 8-14 µm wide at the base.
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Original descripition: Burger, 1980, p. 274
A species of Muderongia, usually flattened dorsoventrally. Pericyst broad, extending into a medium-sized to long tappering horn, two lateral horns and two medium-sized antapical horns, the latter identical in size and shape, conical with pointed tips. Lateral horns not longer than width of body, consisting of a tubiform proximal part and a narrower, tappering, pointed distal part which is bent backwards at varying angles. A small 'elbow' protrusion is often present att bending point where the paracingulum occurs. Endocyst usually displaced towards apex, outline asymmetrical, rhombic to pentagonal with streight or concave sides and rounded to pointed corners, which sometimes protude into lower horn cavities. Both endophragm and periphragm very thin, psilate to punctate. Archaeopyle large, apical (type A/A), with an undulating or zigzag margin indicating six praecingular paraplates; in many specimens an offset parasulcal notch is observed. Operculum free. A paracingulum and posterior parasutures are sometimes seen as faint discontinuos lines on the periphragm.

Emended description: Monteil, 1991, p. 476
Cysts proximate to proximochorate, compressed dorsoventrally, two-layered, cornucavate to delphicavate or circumcavate. Pericyst ceratioid, symmetrical, with 5 prominent tapered horns: 1 apical (axial, type AP l); 2 subequal lateral (bent, type L ll), indented; 2 subequal antapical (not joined, type ATP lll). Sometimes, a "true" elbow may be developed on the postcingular entension (pl. 8, fig. 1-2 in Evitt, 1961 and pl. 1, fig. 6d in Wiseman, 1980). Distal extremities of horns closed and pointed; proximal extremities narrow to wide. Endocyst oval to rhombic, or pentangular, often with protrusions at the base of the horns. Periphragm thin, psilate to scabrate, sometimes locally finely perforated and/or folded; periphragm may be ornamented by processes (stages I to IV). Endophragm thin, psilate to granulate. Archeopyle apical, type (4A), with angular margin. Parasulcal notch marked.
Operculum free. Paratabulation indicated by archeopyle, by low parasutural ridges or septa, and more or less clearly expressed by distribution and type of processes (stages I to IV), when those are present. Occasionally, periphragm showing folds or penitabular microperforations. Formula:?pr, 4', Oa, 6", 6c, 6''', 1p, 1'''',?xs. Pericingulum marked by indentation of lateral horns and by parallel transverse structures. Perisulcus sinuous, marked by parasulcal notch offset.
Dimensions of the holotype (without operculum): 105 µm x 137 µm (overall L x 1, estimated from fig. 9B in Burger, 1980).

Affinities:
Burger, 1980, p. 274: So far this species is unique within the genus in that it develops two fully grown identically antapical horns.
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