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Cleistosphaeridium huguoniotii ssp. pertusum

Cleistosphaeridium huguonioti ssp. pertusum, (Davey, 1969), Lentin and Williams, 1973

NOW Sepispinula pertusa. Originally Cleistosphaeridium huguoniotii var. pertusum, subsequently Cleistosphaeridium huguoniotii subsp. pertusum, thirdly Cleistosphaeridium? huguoniotii subsp. pertusum, fourthly Dapsilidinium ambiguum subsp. pertusum, fifthly Sepispinula huguoniotii subsp. pertusa, sixthly (and now) Sepispinula pertusa.

Holotype: Davey, 1969, pl.7, figs.7
Locus typicus: Fetcham Mill, England
Stratum typicum: Cenomanian

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Original description: [Davey, 1969, p. 156]:

Diagnosis:
A variety of C. huguoniotii possessing spherical to subspherical, smooth walled shell bearing numerous processes. Processes broad-based, tapering distally and terminating with two small recurved spines. Lumen of processes restricted by transverse septa.

Description:
The processes taper distally from a fairly broad base (2.5-4 µm in width) to a narrow neck (c. 0.5 µm in width) before bifurcating to give two short, recurved spines. The processes are hollow but the lumen is traversed by a number of small septa which thus subdivide it, giving the processes a " holey " or vacuolated appearance.
An archaeopyle is only rarely observable.

Affinities:
This variety differs from C. huguoniotii in the form of its processes which are vacuolated, tend to be longer and bear relatively small distal spines. C. huguoniotii found in the same samples possesses processes which are considerably wider distally.
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