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Gonyaulacysta areolata

Gonyaulacysta areolata (Sarjeant, 1961) Lentin and Williams, 1973

Combination illegitimate, since it is a nomenclatural junior synonym of Gonyaulacysta scarburghensis Sarjeant, 1964.
Now Trichodinium scarburghense. Originally Gonyaulax areolata (name illegitimate), subsequently Gonyaulacysta scarburghensis, thirdly Gonyaulacysta areolata (combination illegitimate), fourthly Acanthaulax scarburghensis, fifthly Liesbergia scarburghensis, sixthly (and now) Trichodinium scarburghense. See also Acanthaulax areolata (combination illegitimate).
This species name was originally published as Gonyaulax areolata, which is a validly published junior homonym; it was subsequently given the nom. nov. Gonyaulacysta scarburghensis by Sarjeant, 1964, in accordance with I.C.B.N. Articles 6.4,64.1). This nom. nov. is the oldest legitimate name (I.C.B.N.
Article 6.3). According to Article 55, when a species is transferred, the senior legitimate epithet is transferred to the new generic name.
Tax. sr. synonym of Acanthaulax senta Drugg, 1978, according to Berger, 1986, Jan du Chêne et al., 1986, Riding, 1987, and Kunz, 1990.

Holotype: Sarjeant, 1961, pl.13, fig.13; text-fig.5; Jan du Chêne et al., 1986, pl.3, figs.1-2
Locus typicus: Scarborough Castle Cliff, England
Stratum typicum: Late Callovian-Early Oxfordian

Original diagnosis: Sarjeant, 1961, p.95: Gonyaulax areolata
A species of fossil Gonyaulax with an ovoid theca having the tabulation ?", 1a, 6", 6""", ?1p, 1"""" and with a strong, tapering apical horn rounded at its extremity. The whole thecal surface except the longitudinal and transverse furrows is covered by low ridges forming an irregular reticulate patterning; the thecal wall is very thin but consists of two layers, the outer bearing the ridges. The boundaries of plates and the furrow margins bear low alveolate crests. The longitudinal furrow extends from apex to antapex; it is narrow, as the transverse furrow.

Original description: Sarjeant, 1961, p.96: Gonyaulax areolata
Theca yellowish. Tabulation not easily discerned and a composite text-figure therefore given. Apical tabulation obscure. An anterior intercalary plate clearly present, plus six pre-equatorial plates, of which the first (1") is reduced. Transverse furrow extends around theca in a strong laevo-rotatory spiral such that its two ends differ in antero-posterior position by approximately three times the furrow"s width. Six postequatorial plates certainly present; a posterior intercalary plate appears present but could not be definitely confirmed. No posterior ventral plates present, longitudinal furrow extending to polygonal antapical plate. Surface reticulation of ridges dense; fields delimited by these ridges always small, not regularly polygonal but varying considerably in relative size and shape. Reticulation extending on to apical horn but absent from furrows. Crests consist of linked processes (links sometimes broken) arising from sutures, their points of origin corresponding with junctions of ridges with sutures.
Dimensions: Holotype: Overall length 155 Ám, length of horn 25 Ám, overall breadth 125 Ám; transverse furrow c. 10 Ám wide, longitudinal furrow c. 5 Ám wide. Range: overall length 120-170 Ám, relative proportions being fairly constant.
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