Paragonyaulacysta spinisutura Below, 1990, p.62–64, pl.10, figs.1–6,12–13; text-figs.18a–f.
Holotype: Below, 1990, pl.10, figs.2,5. Contrary to the opinion of Lentin and Williams (1993, p.489), this name is validly published. Below (1987a, p.6) stated that, unless otherwise indicated, all the material is in his personal collection. Since Below (1987a, 1987b, 1990) are parts I to III of a single study, we agree with J. Jansonius (personal communication) that Below's (1987a) repository statement refers to all three publications, fulfilling the requirement of I.C.N. Article 40.7 (see also the discussion on Article 40.7 in the Introduction).
Age: Kimmeridgian.

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Original description (Below, 1990) (translated from German):

Paragonyaulacysta spinisutura n. sp.
Plate 10, Figs. 1-6, 12-13; Fig. 18

Derivation: based on the spines arranged on the fins.
Holotype: Specimen S1-1326(K4)6/8 (Plate 10, Figs. 2, 5).

Description:
Vesicle/plate arrangement of the dinoflagellate Amphiesma cop, pop, cap, 3', 2a, 6'', 7c, 6'", 2'"', as, FM, Is, rs, ps; 1' exsert with adelopore on the suture 1'/6", 1a quadragonal, 2a hexagonal; tegulation 4*, 2**, 4***, 2****, 7c, 4(*), 4(**), 3(***); growth of thecal plates in a gonyaulocoid manner;
Zygote cyst habitus proximochorate, acavate, somewhat elongated polyhedral, epicyst tentoriate with a low apical bulge, somewhat longer than or equal to the highly tentoriate hypocyst, circular to slightly dorsoventrally flattened in equatorial section; wall consisting of a thin pedium and smooth or punctate to evenly reticulated luxuria, numerous, short, cylindrical, evexate, rarely reduced to baculi, spines are arranged finitely in rows, they are isolated or are connected basally by low ridges that extend only along appear as low septa at the cingulum margins, they are rarely found intraareat, the rows of spines divide the cyst into areas; arrangement scheme XPR/cop, pop, 3', 2a, 6", Xc, 6'", 2''", Xs; PR undifferentiated, marked as a group of spines or tiny circular cop surrounded by a horseshoe-shaped pop, cap nonfinate, shadowy; 3 apicals: tiny planate 1' (V) exsert, VIl-nE 2' (VL), VII-nE 3' (VR), caroidale 1a (DL) and VI2a (DDR); 6 precingulate: anterior bigeniculate VI-nE 1'" (VVL), anterior geniculate V-nE 2'' (LVL), anterior geniculate V-nE 3" (DL), anterior linear IV-nE 4" (DDR), Finis 3"/4'' dorsal, anterior geniculate V-nE 5" (RVR), anterior bigeniculate V-nE 6"; Cingulum very deeply incised, left-handed trochospiral with a wide offset of the cingulum beginning and end, thereby the also deeply incised, only partially areate sulcus is sigmoidally curved: as tiny, ps large, omega-shaped, anterior sulcus depressed to 6"; 6 postcingulates: posterior linear IV-nE 1'" (VVL), posterior linear IV-nE 2"' (LVL), posterior geniculate V-nE 3'" (DL), posterior linear IV-nE 4"' (D), posterior linear IV-nE 5"' (R), posterior linear IV-nE 6"' (VVR); 2 antapics: V1"" sloping to VVL, VI2"" approximately pole-side: archaeopyle anterior intercalary, 1a + 2a; Operculum solvate, opercular plates secate, general opercular formula 1a + 2a.

Dimensions of the holotype:
Length 45 µm, width 40 µm.

Note:
In terms of habitus, this proximochorate cyst differs significantly from the previously known proximate species of the genus Paragonyaulagysta, but displays exactly their areation pattern. The habitus, determined by the finately arranged spines, and especially the characteristic punctate surface, initially suggest a relationship to Hystrichodinium species, which are, however, distinguished by different areation and the preecingulate archaeopyle.
Plate overlap is very rare and then only partially reflected as "direct overlap" of areas.

Record:
Specimens 1326, 1334.