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Leberidocysta chlamydata

Leberidocysta chlamydata (Cookson and Eisenack, 1962b, p.496, pl.7, figs.1-3,5-8) Stover and Evitt, 1978, p.60. Emendations: Fechner, 1985, p.119 and Marheinecke, 1992, p.88, both as Leberidocysta chlamydata.

Originally Hexagonifera, subsequently Hexagonifera?, thirdly (and now) Leberidocysta, fourthly Polygonifera.
Mehrotra and Sarjeant, 1984, transferred this species to Polygonifera Habib, 1972. Lentin and Williams, 1985, retained it in Leberidocysta.

Holotype: Cookson and Eisenack, 1962b, pl.7, fig.2; Fensome et al., 1993a, fig.2 - p.1049.
Locus typicus: Brickhouse Bore, Carnarvon, W Australia
Stratum typicum: Albian-Cenomanian

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G.L. Williams short notes on species, Mesozoic-Cenozoic dinocyst course, Urbino, Italy, May 17-22, 1999 - LPP VIEWER CD-ROM 99.5.

Leberidocysta chlamydata (Cookson and Eisenack, 1962b) Stover and Evitt, 1978, emend. Fechner, 1985, emend. Marheinecke, 1992. Emendation from Marheinecke (1992). Cyst habit cavate, consisting of endoblast and periblast. Endoblast ovaloidal with a relatively thick wall (up to 1 µm) composed of pedium and luxuria, which is fused with the pedium to form a foveolate surface. Periblast thin-walled, ovaloidal but usually lying in folds, with a smooth to foveolate surface. There is always contact between periblast and endoblast at the archeopyle margin, and sometimes also in the sulcal region. Tabulation: 4', 0a, 6", Xc, X”’, ?p, ?1"”. Archeopyle apical. Size: overall length 68-75 µm, width 42-54 µm, endocyst 56-68 x 42-54 µm.
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Original description: Cookson and Eisenack, 1962, p. 496: Hexagonifera chlamydata
Shell broadly oval, rather thick-walled, surface finely to coarsely verrucose, completely enclosed in a thin, smooth, hyaline membrane that may extend for some distance laterally and posteriorly or may occasionally be entirely removed.
Dimensions: Holotype: Over-all length 75 Ám; width 76 Ám; shell 60x50 Ám.
Range: Over-all length ca. 68-75 Ám; width ca. 42-54 Ám; shell 56-68x42-54 Ám.

Emended description: Fechner, 1985, p.119
The cavate cyst shows an ellipsoidal relatively thick-walled endophragm with a smooth to granular appearance and a commonly complexly folded thin periphragm. The periphragm and endophragm are attdched only along the Zig-Zag line of the apical archeopyle type tA. In the absence of an archeopyle, endophragm and periphragm are merely loosely attached. While parasutural features of the endophragm are confined to the apical region, expressed simply by an apical horn, the periphragm with its random arrangement of folds provides more information about the paratabulation (Text-fig. 5)
-- Considering the tetragonal apical horn of the endophragm and the hexagonal horn of the periphragm, the existence of 4 - 6 apical paraplates becomes likely. -- The zig-zag line of the archeopyle margin suggests further the presence of 5 - 6 precingular parapIates . -- No signs of intercalary paraplates were found. --Since the antapex of the periphragm is frequently somewhat angular in shape, this area is possibly occupied by only one paraplate. -- Beyond that, the periphragm shows occasionally an arrangement of folds which conceivably mark parasulcus and paracingulum. -- All of the above results in the following preliminary paratabulation formula: 5 - 6", ?C, ?''', ?p, 1'''', ?r.

Affinities: Cookson and Eisenack, 1962, p. 496: Hexagonifera chlamydata
H. chlamydata is distinguishable from the Upper Cretaceous species H. vermiculata by the type of the sulpture and the greater prominence and constancy of the outer membrane.
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