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Biconidinium reductum

Biconidinium reductum, (May, 1980); emend. Kirsch, 1991

Originally Palaeocystodinium, subsequently (and now) Biconidinium.

Holotype: May, 1980, pl.21, fig.20
Locus typicus: Atlantic Highlands, New Jersey
Stratum typicum: Mount Laurel Sand, Campanian-Maastrichtian

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Original description as Palaeocystodinium reductum : [May, 1980, p. 84-85]: (Translation: LPP):

Description:
Periblast spindle-shaped with extended apical and antapical horns; slightly compressed dorso-ventrally, and ovoidal in polar view; periphragm smooth. Horns not as long as endoblast; are pointed to digitate. Antapical horn bears slight deflection similar in position to the accessory spike of P. australinum. Endoblast ovoidal, separated from periblast by continuous lateral and polar pericoels; occupies central bulge of spindle; endophragm smooth.
Nontabulated.
Cingulum formed of low folds in periphragm; especially noticeable in optical cross section; slightly levorotatory less than 1 cingulum width offset. Sulcus barely traceable, formed of slight periphragm depression extending antapically from between terminal ends of cingulum.
Archeopyle not observed.

Dimensions:
Holotype: L x W, 113 x 42 µm; endoblast L x W, 43 x 36 µm; apical horn 27 µm; antapical horn 27 µm; lateral pericoels ca. 4 µm on each side in optical section.
Observed range (4 specimens measured from top of endoblast 27-29 µm; antapical horn measured from bottom of endoblast 24-29 µm; endoblast L x W, 42-43 µm x 35-36 µm. Endophragm less than 1 µm; periphragm less than 1 µm.

Discussion:
The characteristic features of P. reductum are the reduced size and abbreviated horns as compared to P. golzowense Alberti 1961 and P. australinum (C 1965). The presence of cingulum and sulcus, and the absence of an archeopyle are also distinctive. The assignment of this species into Palaeocystodinium is tentative, until further forms are observed which may display the archeopyle type.

Affinities:
P. reductum is similar in appearance to P. ? denticulatum Alberti 1961; however, it lacks the ribbing or striae on the horns, and bears a consistent pericoel about the entire margin of the endoblast. P. australinum and P. golzowense both have much extended horns, giving greater length; whereas P. reductum has relatively abbreviated horns. The apparent lack of an archeopyle (at least the difficulty in detecting it), the size, and the presence of cingulum and sulcus would generally separate D. reductum from P. golzowense and P. australinum.

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Emended description:

Kirsch, 1991, p. 120:

Description:
Spindle-shaped, cavate cyst with usually smooth endo- and periphragm, characterised by an ovoidal central body and one apical and one antapical horn. The smooth to finely granulate endophragm is slightly thicker than the smooth periphragm. Periphragm forms polar processes, developed as long, hollow, distally blunted to conical horns. The central body has no contact to the periphragm and is not involved in archaeopyle forming.
The archeopyle (periarchaeopyle) is apical and indicates tabulation. Archaeopyle margin zigzag.
Tabulation: 4", 6"", ?c, ?""", ?1"""".

Affinities:
The smaller size and the shorter processes distinguish B. reductum from species of Palaeocystodinium.
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