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Muderongia perforata
From Fensome et al., 2019:
Muderongia perforata Alberti, 1961, p.13, pl.2, figs.8–9 (not fig.7 according to Riding et al., 2001, p.28). Emendations: Monteil, 1991b, p.475; Riding et al., 2001, p.28. Holotype: Alberti, 1961, pl.2, fig.8; lost according to Riding et al. (2001, p.28). Lectotype: Alberti, 1961, pl.2, fig.9 and Riding et al., 2001, pl.1, fig.3, designated (as a neotype) by Riding et al. (2001, p.28). Taxonomic senior synonym: Muderongia simplex, according to Jain and Khowaja-Ateequzzaman (1984, p.39) — however, Lentin and Williams (1989, p.252) retained Muderongia perforata. Taxonomic junior synonym: Muderongia pariata, according to Århus in Århus et al. (1990, p.191) — however, Monteil (1991b, p.474) retained Muderongia pariata. Age: Turonian.
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Original description: [Alberti, 1961] (translated from German):
Muderongia perforata n. sp.
Plate 2, Figs. 8 and 9
Holotype: The specimen preserved in preparation no. A 9 and shown on Plate 2, Fig. 8.
Type locality: Pirna borehole (Saxony).
Type stratum: Turonian
Diagnosis: Carapace flattened, sometimes slightly convex, its outline rhombic-oval. With a long, tapering apical horn-like process and two unequal antapical horns, one of which may be largely reduced. Lateral horns strongly projecting, also of unequal length, notched at the free end. Apical and antapical horns blunt at the free end, their points of attachment to the carapace often not lying in the same plane. Membrane of the anterior portion of all horn-like processes coarsely perforated. Always with an inner body.
Additions: A transverse band could not be observed in any specimen. The carapace membrane is delicate and transparent, that of the inner body darker in color.
Relationships: Distinguished from the other species of the genus by the perforation of the anterior portion of the horn-like processes
Holotype dimensions: Length 144 μ, width not measurable as the specimen is in a lateral position. In other specimens, the length varies between 123 μ and 166 μ, the width between 108 μ and 116 μ.
Material: Over 20 specimens.
Occurrence: Turonian of Pirna (relocated?).
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Emended diagnosis: Monteil 1991b, p. 475
Cysts proximate, compressed dorsoventrally, two-layered, circumcavate. Pericyst ceratioid, asymmetrical, with 5 prominent horns: 1 apical (axial, type AP l); 2 subequal lateral (axial, type L 1), indented; 2 inequal antapical (joined, type ATP ll). Left antapical horn longer (up to one and one-half times) than the right one. Horns normal with distal extremities closed and rounded, or open and perforated-scalloped (Pl. 2, fig. 11; Pl. 11, fig. 2); proximal extremities wide. Endocyst oval. Periphragm thin, psilate, perforated in the distal half of the horns; perforations not organized. Endophragm thin, psilate to granulate. Archeopyle apical, type (4A) with angular margin. Parasulcal notch marked. Operculum free. Paratabulation indicated only by archeopyle, by pericingulum and by perisulcus. Formula: ?pr, 4", Oa, 6", s. Pericingulum marked by indentation of lateral horns and by parallel transverse folds. Perisulcus marked by parasulcal notch offset. Right margin of parasulcal area underlined by an oblique fold of periphragm on the left side of the cyst (Pl. 2, fig. 11; Pl. 7, fig.2).
Dimensions of the holotype (with operculum): 144 µm (overall L; I not measurable)
Comparisons: Muderongia perforata emend. differs from M tabulata comb. nov. & emend. in possessing a periphragm perforated in the distal half of horns.
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Muderongia perforata Alberti, 1961, p.13, pl.2, figs.8–9 (not fig.7 according to Riding et al., 2001, p.28). Emendations: Monteil, 1991b, p.475; Riding et al., 2001, p.28. Holotype: Alberti, 1961, pl.2, fig.8; lost according to Riding et al. (2001, p.28). Lectotype: Alberti, 1961, pl.2, fig.9 and Riding et al., 2001, pl.1, fig.3, designated (as a neotype) by Riding et al. (2001, p.28). Taxonomic senior synonym: Muderongia simplex, according to Jain and Khowaja-Ateequzzaman (1984, p.39) — however, Lentin and Williams (1989, p.252) retained Muderongia perforata. Taxonomic junior synonym: Muderongia pariata, according to Århus in Århus et al. (1990, p.191) — however, Monteil (1991b, p.474) retained Muderongia pariata. Age: Turonian.
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Original description: [Alberti, 1961] (translated from German):
Muderongia perforata n. sp.
Plate 2, Figs. 8 and 9
Holotype: The specimen preserved in preparation no. A 9 and shown on Plate 2, Fig. 8.
Type locality: Pirna borehole (Saxony).
Type stratum: Turonian
Diagnosis: Carapace flattened, sometimes slightly convex, its outline rhombic-oval. With a long, tapering apical horn-like process and two unequal antapical horns, one of which may be largely reduced. Lateral horns strongly projecting, also of unequal length, notched at the free end. Apical and antapical horns blunt at the free end, their points of attachment to the carapace often not lying in the same plane. Membrane of the anterior portion of all horn-like processes coarsely perforated. Always with an inner body.
Additions: A transverse band could not be observed in any specimen. The carapace membrane is delicate and transparent, that of the inner body darker in color.
Relationships: Distinguished from the other species of the genus by the perforation of the anterior portion of the horn-like processes
Holotype dimensions: Length 144 μ, width not measurable as the specimen is in a lateral position. In other specimens, the length varies between 123 μ and 166 μ, the width between 108 μ and 116 μ.
Material: Over 20 specimens.
Occurrence: Turonian of Pirna (relocated?).
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Emended diagnosis: Monteil 1991b, p. 475
Cysts proximate, compressed dorsoventrally, two-layered, circumcavate. Pericyst ceratioid, asymmetrical, with 5 prominent horns: 1 apical (axial, type AP l); 2 subequal lateral (axial, type L 1), indented; 2 inequal antapical (joined, type ATP ll). Left antapical horn longer (up to one and one-half times) than the right one. Horns normal with distal extremities closed and rounded, or open and perforated-scalloped (Pl. 2, fig. 11; Pl. 11, fig. 2); proximal extremities wide. Endocyst oval. Periphragm thin, psilate, perforated in the distal half of the horns; perforations not organized. Endophragm thin, psilate to granulate. Archeopyle apical, type (4A) with angular margin. Parasulcal notch marked. Operculum free. Paratabulation indicated only by archeopyle, by pericingulum and by perisulcus. Formula: ?pr, 4", Oa, 6", s. Pericingulum marked by indentation of lateral horns and by parallel transverse folds. Perisulcus marked by parasulcal notch offset. Right margin of parasulcal area underlined by an oblique fold of periphragm on the left side of the cyst (Pl. 2, fig. 11; Pl. 7, fig.2).
Dimensions of the holotype (with operculum): 144 µm (overall L; I not measurable)
Comparisons: Muderongia perforata emend. differs from M tabulata comb. nov. & emend. in possessing a periphragm perforated in the distal half of horns.
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