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Polygonifera aspera
Polygonifera? aspera Dodekova, 1990
Dodekova, 1990 questionably included this species in Polygonifera.
Holotype: Dodekova, 1990, pl.4, fig.13
Locus typicus: borehole R-l Hrabrovo, depth 1977.2 m, Sultanci Formation, East Bulgaria
Stratum typicum: Middle Callovian
Original diagnosis: Dodekova, 1990, p.41
Cyst camocavate with spherical to subspherical endocyst and polyedrical pericyst, lacking apical and antapical horn. Endophragm thick, scabrate to irregularly reticulate in one and the same specimen, with single rare tuberculae. Periphragm very thin, readily being deformed. Its surface is scabrate to granulate or with very fine indistinct reticulation. There are traces of tabulation on it marked by vague low ribs. Tabulation formula: 1-2?pr, 4", X?a, 6", X?c, 6""", 1p, 1pv+Xs, 1"""". Archeopyle combination, apicalintercalary, type 4Atl/4AtT. Operculum free, composed of 4 apical plates. The intercalary plates are falling apart together with the operculum, or later one by one (pl. IV, figs 11, 12).
Dimensions: Holotype: diameter of endocyst 64 µm, diameter of pericyst 79 µm. Range: diameter of endocyst 52-64 µm, diameter of pericyst 64-86 µm (22 specimens measured).
Description and remarks: Dodekova, 1990, p.41
The new species here introduced is highly vulnerable to secondary deformations. Hence, the great variability in its shape. Similar variation in the shape and cavation of P. eisenackii Merhotra and Sarjeant, 1984 is mentioned by Merhotra and Sarjeant (1984, p. 46). 1n P.? aspera the two wall layers may be closely stuck together or widely separated, no matter which part of the cyst is considered. The two layers are always appressed on the apex forming thus a faint broadlyconical swelling. Antapical pericoel may be present, but is commonly lacking. When present it lacks the shape typical for the genus Polygonifera (Habib, 1972, pl.5, figs.1-2; Brenner, 1988, pl.12, figs 3-6), but is largely rounded. The typical for the genus ventral, antapical and lateral pericoels are, in the new species, rarely, as if occasionally manifested. Same is the case with the dorso-ventral flattening pronounced in the genus Polvgonifera. The well-preserved specimens of P.? aspera have shperical or subspherical endocyst. Regardless of the thick endophtagm the epiendocyst is being often deformed, disintegrating at? plates or dismembering. Endosyst's ornamentation is irregularly distributed. Usually the endosulcus is poorly ornamented, slightly concave, always having flagellar mark (pl. IV, figs.13,16). Tabulation indicated on the periphragm by low ribs, hardly distinguishable for the numerous secondary folds. Distinct cingulum and two ventral hypocyst lobes, characteristic for the genus, are sometimes observed. They are far poorly expressed in comparision with those of the type species.
The archeopyle is apical-intercalary concerning both the endo-and periphragm, which are in this part of the cyst closely appressed. Type--4AtI/4AtI. Initially 4 apical plates fall away. Specimens with their peri-endoapical operculum half open or slightly displaced are found (pl. IV, fig., 1-2). A vast amount of such free operculums are available in the slides (pl. IV, fig.12). Subsequently or simultaneously the Xa intercalary plates drop off. Specimens with 3a symmetrically positioned, detached intercalary plates around the apical archeopyle are observed (pl. IV, fig.3). In some free apical operculums half-detached intercalry plates are available (pl. IV, fig.11). Finally, some specimens have widely opened archeopyle (pl. IV, fig.16). All the aforementioned suggests the presence of series of intercalary plates around the plates of the apical series. Anterior intercalary plates in genus Polygonifera have hithertonotbeen reported. The latter and the mentioned earlier morphological differences, as well as certain features of the new speciesP.? aspera similar to those of the type species P. evittii gave us the ground to be referring with great reservations the species P.? aspera to the genus Polygonifera.
Dodekova, 1990 questionably included this species in Polygonifera.
Holotype: Dodekova, 1990, pl.4, fig.13
Locus typicus: borehole R-l Hrabrovo, depth 1977.2 m, Sultanci Formation, East Bulgaria
Stratum typicum: Middle Callovian
Original diagnosis: Dodekova, 1990, p.41
Cyst camocavate with spherical to subspherical endocyst and polyedrical pericyst, lacking apical and antapical horn. Endophragm thick, scabrate to irregularly reticulate in one and the same specimen, with single rare tuberculae. Periphragm very thin, readily being deformed. Its surface is scabrate to granulate or with very fine indistinct reticulation. There are traces of tabulation on it marked by vague low ribs. Tabulation formula: 1-2?pr, 4", X?a, 6", X?c, 6""", 1p, 1pv+Xs, 1"""". Archeopyle combination, apicalintercalary, type 4Atl/4AtT. Operculum free, composed of 4 apical plates. The intercalary plates are falling apart together with the operculum, or later one by one (pl. IV, figs 11, 12).
Dimensions: Holotype: diameter of endocyst 64 µm, diameter of pericyst 79 µm. Range: diameter of endocyst 52-64 µm, diameter of pericyst 64-86 µm (22 specimens measured).
Description and remarks: Dodekova, 1990, p.41
The new species here introduced is highly vulnerable to secondary deformations. Hence, the great variability in its shape. Similar variation in the shape and cavation of P. eisenackii Merhotra and Sarjeant, 1984 is mentioned by Merhotra and Sarjeant (1984, p. 46). 1n P.? aspera the two wall layers may be closely stuck together or widely separated, no matter which part of the cyst is considered. The two layers are always appressed on the apex forming thus a faint broadlyconical swelling. Antapical pericoel may be present, but is commonly lacking. When present it lacks the shape typical for the genus Polygonifera (Habib, 1972, pl.5, figs.1-2; Brenner, 1988, pl.12, figs 3-6), but is largely rounded. The typical for the genus ventral, antapical and lateral pericoels are, in the new species, rarely, as if occasionally manifested. Same is the case with the dorso-ventral flattening pronounced in the genus Polvgonifera. The well-preserved specimens of P.? aspera have shperical or subspherical endocyst. Regardless of the thick endophtagm the epiendocyst is being often deformed, disintegrating at? plates or dismembering. Endosyst's ornamentation is irregularly distributed. Usually the endosulcus is poorly ornamented, slightly concave, always having flagellar mark (pl. IV, figs.13,16). Tabulation indicated on the periphragm by low ribs, hardly distinguishable for the numerous secondary folds. Distinct cingulum and two ventral hypocyst lobes, characteristic for the genus, are sometimes observed. They are far poorly expressed in comparision with those of the type species.
The archeopyle is apical-intercalary concerning both the endo-and periphragm, which are in this part of the cyst closely appressed. Type--4AtI/4AtI. Initially 4 apical plates fall away. Specimens with their peri-endoapical operculum half open or slightly displaced are found (pl. IV, fig., 1-2). A vast amount of such free operculums are available in the slides (pl. IV, fig.12). Subsequently or simultaneously the Xa intercalary plates drop off. Specimens with 3a symmetrically positioned, detached intercalary plates around the apical archeopyle are observed (pl. IV, fig.3). In some free apical operculums half-detached intercalry plates are available (pl. IV, fig.11). Finally, some specimens have widely opened archeopyle (pl. IV, fig.16). All the aforementioned suggests the presence of series of intercalary plates around the plates of the apical series. Anterior intercalary plates in genus Polygonifera have hithertonotbeen reported. The latter and the mentioned earlier morphological differences, as well as certain features of the new speciesP.? aspera similar to those of the type species P. evittii gave us the ground to be referring with great reservations the species P.? aspera to the genus Polygonifera.