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Sumatradinium druggii

Sumatradinium druggii Lentin et al., 1994

Holotype: Lentin et al., 1994, pl.1, fig.4
Paratypes; Lentin et al., 1994, pl.1, figs.5,7
Locus typicus: Evangeline H-98 well, Scotian Shelf, offshore Eastern Canada
Stratum typicum: Miocene (Tortonian)

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G.L. Williams short notes on species, Mesozoic-Cenozoic dinocyst course, Urbino, Italy, May 17-22, 1999

Sumatradinium druggii Lentin et al., 1994. Sumatradinium druggii has processes of variable length and morphology, which usually have annular thickenings and closed furcate terminations. In some processes the “annular thickenings” may be septa, but this is impossible to tell under the light microscope. Size: body length 73-98 µm, width 64-86 µm, minimum length of processes 6-14 µm, maximum length of processes 10-18 µm.
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Orignal diagnosis: Lentin et al., 1994,p. 574:
A species of Sumatradinium with hollow, distally closed processes with typically multifurcate terminations; the processes usually have annular thickenings.

Description:
Shape: Cyst weakly to strongly dorsoventrally compressed, ambitus subcircular, suboval, rounded pentagonal to peridinioid. Apex rounded; antapex rounded to weakly bilobate and symmetrical or asymmetrical. Epicyst and hypocyst appear to be more or less equal in size.
Wall relationships: Usually acavate with autophragm only or with endophragm and periphragm appressed; rarely circumcavate, with periphragm and endophragm narrowly separated.
Wall features: Surface ornamentation comprises a complete to incomplete reticulum or, locally, a granulation; the ornament may extend onto lower part of processes. Processes variable in length and width, especially between specimens; hollow, generally with annular thickenings along their length, tubular or tapering, usually distally closed, possibly occasionally open, with multifurcate extremities. Some furcations may be secondarily branched. Mode of excystment: Archeopyle usually involves the loss of a large hexa isodeltaform middorsal intercalary paraplate, presumably 2a, posteriorly extending almost to the equator of the cyst. The operculum is usually free but may be adnate anteriorly. Rarely, accessory archeopyle sutures develop between paraplates adjacent to 2a and a second intercalary paraplate may be dislodged. When two wall layers are observed, they appear fused on the operculum. Paratabulation: Generally indicated by archeopyle only and assumed to be peridinialean; occasionally the alignment of processes suggests a paracingulum and possibly other parasutures. Parasulcus not identified. Size: Holotype: body length 76 µm, body width 70 µm, length of processes up to 15 µm. Range: body length 73(81)98 µm (19 specimens), body width 64(71)86 µm (19 specimens), minimum length of processes 6(10)14 µm (17 specimens), maximum length of processes 10(13)18 µm (17 specimens).

Affinities:
Lentin et al., 1994, p.574: The genus Sumatradinium includes four species: S. hispidum (Drugg, 1970) Lentin and Williams, 1976 (the type), S. druggii sp.nov., S. pustulosum sp.nov., and S. soucouyantiae de Verteuil and Norris, 1992. The species can be readily separated by the nature of the processes.
Sumatradinium hispidum has short (up to 5 µm), closed, spine-like processes, which are acuminate distally and commonly penitabulate or possibly sutural. Sumatradinium soucouyantiae has processes 6--20 Ám in length, which are usually hollow and distally acuminate to rounded or clavate, not furcate, and always closed. Sumatradinium druggii has processes of variable length and morphology, which usually have annular thickenings and closed furcate terminations. In some processes the "annular thickenings" may be septa but this is impossible to prove under the light microscope. Sumatradinium pustulosum has a low, pustulate ornamentation.
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