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Cassiculosphaeridia intermedia
From Fensome et al., 2019:
Cassiculosphaeridia intermedia Slimani, 1994, p.99–100, pl.18, figs.13–15. Holotype: Slimani, 1994, pl.18, figs.14–15. Questionable
assignment: Slimani (1994, p.99). Taxonomic junior synonym: Chytroeisphaeridia everricula (name not validly
published), according to Slimani (1994, p.99). Age: late Campanian–Danian.
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Original description (Slimani, 1994) Translated PKB 2024
Cassiculosphaeridia? intermedia sp. nov. (P1.18, Fig.13-15)
Latin: intermedins, intermediate; reference to this species which is intermediate between C. pygmaea and C. magna.
Holotype: Turnhout -892.87 m, preparation 7, coord. E.F. W27/1.
Paratvpe: Beutenaken ech. 16, preparation 6, coord. E.F. N43.
Type locality: Turnhout -892.87 m. Typical horizon: Lower Maastrichtian. Synonymy: 1974: Chytroeisphaeridia everricula sp. nov. in WILSON, p.221-223, pl. 15, fie. 10-12, text-fig. 36. 1983a: Chytroeisphaeridia everricula in WILSON - FOUCHER, pl. 6, fig. 6-7. 1985: Chytroeisphaeridia everricula in WILSON - FOUCHER in ROBASZYNSKI et al., pl. 12, fig. 7-8. 1991: Cassiculosphaeridia sp. A in LOUWyE, p. 90, P 1.2, fig. 9. 1 99 1 Chytroeisphaeridia sp. A KIRSCH, p. 52, pl.30, fig. 1. 1993: "Chytroeisphaeridia everricula" in WILSON, 1974 - SCHIOLER and WILSON, pl. 1, fig. 7.
Diagnosis: Large, proximate, subovoidal to subspheroidal cyst with an autophragm that bears an irregular reticulum. The archaeopyle is apical with an operculum often adnate. Dimensions: Holotype: length: 66 pm, width: 64 pm.
Variations: length: 54 - 70, width: 43 - 64 pm. Number of specimens measured: 20.
Material: > 120 copies. Description: The autophragm, 1 to 1.5 µm thick, is smooth to densely dotted. The reticulum is irregular and low (1 to 1.5 pm in height). The ridges of the reticulum are solid, smooth or dotted, and generally thicker at the points of their junctions; the meshes have a rounded to slightly angular outline, and a diameter which varies from 2 to 8 pm with an average of 4 to 5 pm. The archaeopyle is type (tA) apical, wide (36 pm in Phototype), with a zig-zag margin and more or less developed accessory sutures. The operculum, more or less angular in outline, is often attached. Paratabulation is indicated only by sutures of the archaeopyle: 74', 5-76". The paracingulum and parasulcus are not indicated.
Note: The reticulate autophragm and the apical archaeopyle with a zig-zag margin in this species allow us to classify it in the genus Cassiculosphaeridia DAVEY (1969a), but this attribution is provisional because the operculum is apparently attached. Comparison: Referring especially to the reticulum, it appears that the present species corresponds to the specimens of Chytroeisphaeridia everricula, figured by WILSON (1974) in his Ph. D. work, by FOUCHER (1983a) and BY FOUCHER in ROBASZVNSKI el al. (1985). The specimens called Cassiculosphaeridia sp. A in LOUWYE (1991) could also correspond to our species. Cassiculosphaeridia pygmaea STEVENS (1987) (Basal Berriasian, Australia) has the same appearance as Cassiculosphaeridia? intermedia sp. nov but it differs by its smaller size, its ellipsoidal shape and by its reticulum with smaller meshes. Australian specimens determined as Cassiculosphaeridia magna DAVY (1974) by STOVER and HELBY (1987c: fig. 20, E-L), (Lower Cretaceous) and by BACKHOUSE (1988: p. 78, pl. 20, fig. 5-7) (Valanginian - Aptian) appear to have a reticulum whose meshes are almost comparable in diameter with those of the species studied, but they are distinguished by the autophragm and the walls which are thicker, as well as by the subspheroidal shape and the larger size of the cyst. The original C. magna material (Berriasian, Britain) has a broader reticle, with thicker, much taller (4-9 pm), process-supported ridges. Referring to the size, shape and reticle of our specimens, we suggest that Cassiculosphaeridia? intermedia sp. nov. is an intermediate form between C. pygmaea and C. magna.
Stratigraphic distribution: Beutenaken: sample 2 to 21, Halembaye: ex. 12 to 87; Upper Campanian - Maastrichtian upper. (B. mucronata zone - B. junior zone). Turnhout: -966.50 to -739.31 m. Upper Campanian - Danien. In WILSON (1974), Campanian sup. - Maastrichtian upper. (langei Zone - casimirovensis and junior Zone), Denmark; Upper Maastrichtian (junior Zone), Maastricht region. FOUCHER (1983a), and FOUCHER in ROBASZYNSKl et al. (1985), Campanian - Upper Maastrichtian, Maastricht region. In LOUWYE (1991), Campanian. Knokke. Belgium. KIRSCH (1991), Maastrichtian, Germany. SCHIOLER and WILSON (1993), Maastrichtian. North Sea, Danish territorial area.
Cassiculosphaeridia intermedia Slimani, 1994, p.99–100, pl.18, figs.13–15. Holotype: Slimani, 1994, pl.18, figs.14–15. Questionable
assignment: Slimani (1994, p.99). Taxonomic junior synonym: Chytroeisphaeridia everricula (name not validly
published), according to Slimani (1994, p.99). Age: late Campanian–Danian.
---------------------------------------------------------------------------------------------------
Original description (Slimani, 1994) Translated PKB 2024
Cassiculosphaeridia? intermedia sp. nov. (P1.18, Fig.13-15)
Latin: intermedins, intermediate; reference to this species which is intermediate between C. pygmaea and C. magna.
Holotype: Turnhout -892.87 m, preparation 7, coord. E.F. W27/1.
Paratvpe: Beutenaken ech. 16, preparation 6, coord. E.F. N43.
Type locality: Turnhout -892.87 m. Typical horizon: Lower Maastrichtian. Synonymy: 1974: Chytroeisphaeridia everricula sp. nov. in WILSON, p.221-223, pl. 15, fie. 10-12, text-fig. 36. 1983a: Chytroeisphaeridia everricula in WILSON - FOUCHER, pl. 6, fig. 6-7. 1985: Chytroeisphaeridia everricula in WILSON - FOUCHER in ROBASZYNSKI et al., pl. 12, fig. 7-8. 1991: Cassiculosphaeridia sp. A in LOUWyE, p. 90, P 1.2, fig. 9. 1 99 1 Chytroeisphaeridia sp. A KIRSCH, p. 52, pl.30, fig. 1. 1993: "Chytroeisphaeridia everricula" in WILSON, 1974 - SCHIOLER and WILSON, pl. 1, fig. 7.
Diagnosis: Large, proximate, subovoidal to subspheroidal cyst with an autophragm that bears an irregular reticulum. The archaeopyle is apical with an operculum often adnate. Dimensions: Holotype: length: 66 pm, width: 64 pm.
Variations: length: 54 - 70, width: 43 - 64 pm. Number of specimens measured: 20.
Material: > 120 copies. Description: The autophragm, 1 to 1.5 µm thick, is smooth to densely dotted. The reticulum is irregular and low (1 to 1.5 pm in height). The ridges of the reticulum are solid, smooth or dotted, and generally thicker at the points of their junctions; the meshes have a rounded to slightly angular outline, and a diameter which varies from 2 to 8 pm with an average of 4 to 5 pm. The archaeopyle is type (tA) apical, wide (36 pm in Phototype), with a zig-zag margin and more or less developed accessory sutures. The operculum, more or less angular in outline, is often attached. Paratabulation is indicated only by sutures of the archaeopyle: 74', 5-76". The paracingulum and parasulcus are not indicated.
Note: The reticulate autophragm and the apical archaeopyle with a zig-zag margin in this species allow us to classify it in the genus Cassiculosphaeridia DAVEY (1969a), but this attribution is provisional because the operculum is apparently attached. Comparison: Referring especially to the reticulum, it appears that the present species corresponds to the specimens of Chytroeisphaeridia everricula, figured by WILSON (1974) in his Ph. D. work, by FOUCHER (1983a) and BY FOUCHER in ROBASZVNSKI el al. (1985). The specimens called Cassiculosphaeridia sp. A in LOUWYE (1991) could also correspond to our species. Cassiculosphaeridia pygmaea STEVENS (1987) (Basal Berriasian, Australia) has the same appearance as Cassiculosphaeridia? intermedia sp. nov but it differs by its smaller size, its ellipsoidal shape and by its reticulum with smaller meshes. Australian specimens determined as Cassiculosphaeridia magna DAVY (1974) by STOVER and HELBY (1987c: fig. 20, E-L), (Lower Cretaceous) and by BACKHOUSE (1988: p. 78, pl. 20, fig. 5-7) (Valanginian - Aptian) appear to have a reticulum whose meshes are almost comparable in diameter with those of the species studied, but they are distinguished by the autophragm and the walls which are thicker, as well as by the subspheroidal shape and the larger size of the cyst. The original C. magna material (Berriasian, Britain) has a broader reticle, with thicker, much taller (4-9 pm), process-supported ridges. Referring to the size, shape and reticle of our specimens, we suggest that Cassiculosphaeridia? intermedia sp. nov. is an intermediate form between C. pygmaea and C. magna.
Stratigraphic distribution: Beutenaken: sample 2 to 21, Halembaye: ex. 12 to 87; Upper Campanian - Maastrichtian upper. (B. mucronata zone - B. junior zone). Turnhout: -966.50 to -739.31 m. Upper Campanian - Danien. In WILSON (1974), Campanian sup. - Maastrichtian upper. (langei Zone - casimirovensis and junior Zone), Denmark; Upper Maastrichtian (junior Zone), Maastricht region. FOUCHER (1983a), and FOUCHER in ROBASZYNSKl et al. (1985), Campanian - Upper Maastrichtian, Maastricht region. In LOUWYE (1991), Campanian. Knokke. Belgium. KIRSCH (1991), Maastrichtian, Germany. SCHIOLER and WILSON (1993), Maastrichtian. North Sea, Danish territorial area.