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Flandrecysta furcata
From Fensome et al., 2019:
Flandrecysta furcata Slimani, 1994, p.53–55, pl.8, figs.15–18; text-figs.6A–B. Holotype: Slimani, 1994, pl.8, figs.15–18; textfigs.
6A–B. Age: early Campanian–early Maastrichtian.
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Slimani (1994): Original diagnosis (translation PKB 2025):
Name derivation: Latin: furcatus-à-um: forked; reference to the furcate processes.
Holiday: Turnhout -933 m, preparation 12, E.F. coord. U50/3.
Type locality: Turnhout -933 m.
Type horizon: Campanian.
Synonyms: 1992: Hystrichostrogylon memhraniphorum AGELOPOULOS, 1964 - MARHEINECKE, p. 37, s. 5, fig. 1.
Diagnosis: Flandrecysta cyst characterized by the presence of long, hollow dorsal paracingulate and postcingulate processes that are distally closed. There, they are closed as well as proximally.
Dimensions: Holotype: endocyst, length: 44 µm, width: 33 µm; pericyst, length: 70 µm, width: 50 µm. Variations: endocyst, length: 32-40 µm, width: 24-39 µm; pericyst, length: 40-50 µm, width: 60-80 µm. Process length: 8-20 µm. Ventral opening diameter: approximately 40 µm. Number of specimens measured: 11.
Material: > 40 specimens.
Description: Skolochorate cyst; the smooth endocyst is ovoidal to ellipsoidal, often bearing an apical bump; the smooth to slightly scabrous pericyst has a subhemicircular outline. The processes are longer where the two membranes are joined. The hypocyst often bears 4 long, hollow postcingulate processes, closed proximally and distally, without proximal or distal connections and bifurcated to polyfurcated. On the epicyst, the pericyst bears short, acuminate or bifid gonal and intergonal processes connected proximally by lines or low ridges that indicate a paratabulation whose formula is: 4', 6", ?6c. 4"',. The paracingulum is marked by two parallel lines and by gonal and intergonal processes, the longest of which are forked and located dorsally where the periphragm and endophragm are joined. A large part of the parasulcus is occupied by the ventral opening of the periphragm. The archaeopyle is precingulate of type P(3"), the operculum is free or recovered inside the cyst.
Comparison: Flandrecysta furcata sp. nov. is characterized by its Achomosphaera-like dorsal paracingulate and postcingulate processes. The presence of processes differentiates the species from Flandrecysta inflata (HEILLMANN-CLAUSEN in THOMSEN and HEILMANN-CLAUSEN, 1985) comb.nov. The type material of Hystrichostrogylon membraniphorum AGELOPOULOS (1964) or others such as the specimens encountered in Great Britain and named Achomosphaera membraniphora (AGELOPOULOS, 1964) by EATON (1976: p. 237-238, pl. 5, fig. 8-10, text-fig. 5) and those figured by FECHNER (1985: pl. 2, fig. 1,2,3,6, text-fig. 4), also present Achomosphaera-type processes but the longest are located in the apical region, the pericoel is more developed there especially in the antapical region, and the short processes on the pericoel are strongly forked instead of being acuminate or bifid as in the present species; these characters clearly separate them from the genus studied. The specimen shown under the name of Hystrichostrogylon memhraniferum by MARHEINECKE (1992: pl. 5, fig. 1). presents the same characters as Flandrecystafurcata sp. nov.
Flandrecysta furcata Slimani, 1994, p.53–55, pl.8, figs.15–18; text-figs.6A–B. Holotype: Slimani, 1994, pl.8, figs.15–18; textfigs.
6A–B. Age: early Campanian–early Maastrichtian.
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Slimani (1994): Original diagnosis (translation PKB 2025):
Name derivation: Latin: furcatus-à-um: forked; reference to the furcate processes.
Holiday: Turnhout -933 m, preparation 12, E.F. coord. U50/3.
Type locality: Turnhout -933 m.
Type horizon: Campanian.
Synonyms: 1992: Hystrichostrogylon memhraniphorum AGELOPOULOS, 1964 - MARHEINECKE, p. 37, s. 5, fig. 1.
Diagnosis: Flandrecysta cyst characterized by the presence of long, hollow dorsal paracingulate and postcingulate processes that are distally closed. There, they are closed as well as proximally.
Dimensions: Holotype: endocyst, length: 44 µm, width: 33 µm; pericyst, length: 70 µm, width: 50 µm. Variations: endocyst, length: 32-40 µm, width: 24-39 µm; pericyst, length: 40-50 µm, width: 60-80 µm. Process length: 8-20 µm. Ventral opening diameter: approximately 40 µm. Number of specimens measured: 11.
Material: > 40 specimens.
Description: Skolochorate cyst; the smooth endocyst is ovoidal to ellipsoidal, often bearing an apical bump; the smooth to slightly scabrous pericyst has a subhemicircular outline. The processes are longer where the two membranes are joined. The hypocyst often bears 4 long, hollow postcingulate processes, closed proximally and distally, without proximal or distal connections and bifurcated to polyfurcated. On the epicyst, the pericyst bears short, acuminate or bifid gonal and intergonal processes connected proximally by lines or low ridges that indicate a paratabulation whose formula is: 4', 6", ?6c. 4"',. The paracingulum is marked by two parallel lines and by gonal and intergonal processes, the longest of which are forked and located dorsally where the periphragm and endophragm are joined. A large part of the parasulcus is occupied by the ventral opening of the periphragm. The archaeopyle is precingulate of type P(3"), the operculum is free or recovered inside the cyst.
Comparison: Flandrecysta furcata sp. nov. is characterized by its Achomosphaera-like dorsal paracingulate and postcingulate processes. The presence of processes differentiates the species from Flandrecysta inflata (HEILLMANN-CLAUSEN in THOMSEN and HEILMANN-CLAUSEN, 1985) comb.nov. The type material of Hystrichostrogylon membraniphorum AGELOPOULOS (1964) or others such as the specimens encountered in Great Britain and named Achomosphaera membraniphora (AGELOPOULOS, 1964) by EATON (1976: p. 237-238, pl. 5, fig. 8-10, text-fig. 5) and those figured by FECHNER (1985: pl. 2, fig. 1,2,3,6, text-fig. 4), also present Achomosphaera-type processes but the longest are located in the apical region, the pericoel is more developed there especially in the antapical region, and the short processes on the pericoel are strongly forked instead of being acuminate or bifid as in the present species; these characters clearly separate them from the genus studied. The specimen shown under the name of Hystrichostrogylon memhraniferum by MARHEINECKE (1992: pl. 5, fig. 1). presents the same characters as Flandrecystafurcata sp. nov.