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Enneadocysta harrisii
Enneadocysta harrisii Stover and Williams, 1995
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Stover and Williams, 1995, p. 111-112
Plate 3, Fiures 2a-c, 3, 4a-d, 5; text-figure 1, nos. D, H
Cordosphaeridium capricornum Cookson and Eisenack 1965, pl. 15, figs. 1, 9 only [not fig. 7 which is the holotype of Cordosphaeridium capricornum.
Description:
Shape: Cysts skolochorate with subspherical to nearly subspherical central body.
Wall relationships: Autophragm only.
Wall features: Autophragm faintly granular to pitted: central body typically bearing 20 intratabular processes, 11 on epicyst and nine on hypocyst; paracingular processes rarely present. Process stems solid, smooth or fibroid, expanded slightly proximally, usually more so distally, and may be split longitudinally or fenestrate. Stems vary in width and those on dorsal and lateral processes usually wider than those on ventral processes. Lengths of stems fairly constant on individual specimens. Most process tips licrate, others simply or complexly branched. Licrate tips have weakly to strongly arcuate terminations with smooth inner margins and finely denticulate outer margins. Processes on paraplates 2"" to 5"" have licrite tips, those on 1"" and anterior sulcal have branched endings and the process on paraplate 6"" may have licrite or branched tips. Ends of arcuations on processes with licrite tips directed equatorially. On hypocyst, the dorsal, lateral and antapical processes have licrate tips; those on postcingular processes have the ends of the arcuations directed equatorially and those on the opposed antapical processes face each other medially. On some specimens, the dorsal ends of antapical processes are joined by a narrow trabecula. Distal end of the posterior sulcal process branched or expanded.
Excystment type: Archeopyle apical; operculum tetratabular, simple, equidimensional or slightly longer transversely than dorso-ventrally and free.
Paratabulation: Indicated on epicyst by principal and short accessory archeopyle sutures and by positions of the intratabular processes; on hypocysts solely by process locations. Process formula: 4", 6"", 0-?1c, 6""", 2"""", 1as, 1ps. Seven processes present subjacent to archeopyle margin. Last precingular paraplate is wider than first and parasulcal notch is medial or offset slightly to the left. Hypocystal processes arranged in partiform pattern.
Paracingulum: Indicated usually by the absence of processes.
Parasulcus: Indicated anteriorly by sulcal notch and anterior parasulcal process and posteriorly by posterior parasulcal process.
Size: Intermediate to large; measurements of 10 specimens: body diameter 43 Ám (50 Ám) 68 Ám, overall diameter 75 Ám (83 Ám) 103 Ám, process length 12 Ám (22 Áum) 30 Ám - most more than 20 Ám long, process width 2 Ám to 14 Ám - most less than 10 Ám wide at mid-height, widths of process tips 9 Ám (23 Ám) 30 Ám. Mean values shown in parentheses.
Comments:
The number of processes is essentially consistent on specimens of Enneadocysta harrisii. Of the 44 specimens examined all had six precingular processes, one anterior parasulcal process and nine hypocystal processes (text-fig. 1, no. H). One specimen had an extra narrow, ventral, presumably equatorial, process that could be paracingular, otherwise cingular processes are absent (pl. 3, figs. 4a-c). Also on two specimens the branched tips of processes 1"" and as joined those of processes 1"" and posterior sulcal, respectively. The greatest variation is in the width of the processes at midlength and in the width of the distal expansions.
Holotype:
Specimen illustrated in Cookson and Eisenack (1965, pl. 15, fig. 1). Location: Palaeontological collection, National Museum of Victoria, Melbourne, Victoria, Australia.
Type locality and stratum:
Locality: Browns Creek outwash section (first gully west of Browns Creek) about 0.4 km east of the mouth of the Johannas River. Aire District, Victoria, Australia.
Stratum: Upper Eocene (Priabonian) carbonaceous clay about 13.1 meters above Notostrea Greensand, Browns Creek Clays (sample E of Cookson and Eisenack 1965).
Stratigraphic and geographic data:
Late Eocene; the species is known only from the type locality, Browns Creek, Victoria, Australia and occurs between 1.8 meters below to 13.1 meters above the Notostrea Greensand.
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Stover and Williams, 1995, p. 111-112
Plate 3, Fiures 2a-c, 3, 4a-d, 5; text-figure 1, nos. D, H
Cordosphaeridium capricornum Cookson and Eisenack 1965, pl. 15, figs. 1, 9 only [not fig. 7 which is the holotype of Cordosphaeridium capricornum.
Description:
Shape: Cysts skolochorate with subspherical to nearly subspherical central body.
Wall relationships: Autophragm only.
Wall features: Autophragm faintly granular to pitted: central body typically bearing 20 intratabular processes, 11 on epicyst and nine on hypocyst; paracingular processes rarely present. Process stems solid, smooth or fibroid, expanded slightly proximally, usually more so distally, and may be split longitudinally or fenestrate. Stems vary in width and those on dorsal and lateral processes usually wider than those on ventral processes. Lengths of stems fairly constant on individual specimens. Most process tips licrate, others simply or complexly branched. Licrate tips have weakly to strongly arcuate terminations with smooth inner margins and finely denticulate outer margins. Processes on paraplates 2"" to 5"" have licrite tips, those on 1"" and anterior sulcal have branched endings and the process on paraplate 6"" may have licrite or branched tips. Ends of arcuations on processes with licrite tips directed equatorially. On hypocyst, the dorsal, lateral and antapical processes have licrate tips; those on postcingular processes have the ends of the arcuations directed equatorially and those on the opposed antapical processes face each other medially. On some specimens, the dorsal ends of antapical processes are joined by a narrow trabecula. Distal end of the posterior sulcal process branched or expanded.
Excystment type: Archeopyle apical; operculum tetratabular, simple, equidimensional or slightly longer transversely than dorso-ventrally and free.
Paratabulation: Indicated on epicyst by principal and short accessory archeopyle sutures and by positions of the intratabular processes; on hypocysts solely by process locations. Process formula: 4", 6"", 0-?1c, 6""", 2"""", 1as, 1ps. Seven processes present subjacent to archeopyle margin. Last precingular paraplate is wider than first and parasulcal notch is medial or offset slightly to the left. Hypocystal processes arranged in partiform pattern.
Paracingulum: Indicated usually by the absence of processes.
Parasulcus: Indicated anteriorly by sulcal notch and anterior parasulcal process and posteriorly by posterior parasulcal process.
Size: Intermediate to large; measurements of 10 specimens: body diameter 43 Ám (50 Ám) 68 Ám, overall diameter 75 Ám (83 Ám) 103 Ám, process length 12 Ám (22 Áum) 30 Ám - most more than 20 Ám long, process width 2 Ám to 14 Ám - most less than 10 Ám wide at mid-height, widths of process tips 9 Ám (23 Ám) 30 Ám. Mean values shown in parentheses.
Comments:
The number of processes is essentially consistent on specimens of Enneadocysta harrisii. Of the 44 specimens examined all had six precingular processes, one anterior parasulcal process and nine hypocystal processes (text-fig. 1, no. H). One specimen had an extra narrow, ventral, presumably equatorial, process that could be paracingular, otherwise cingular processes are absent (pl. 3, figs. 4a-c). Also on two specimens the branched tips of processes 1"" and as joined those of processes 1"" and posterior sulcal, respectively. The greatest variation is in the width of the processes at midlength and in the width of the distal expansions.
Holotype:
Specimen illustrated in Cookson and Eisenack (1965, pl. 15, fig. 1). Location: Palaeontological collection, National Museum of Victoria, Melbourne, Victoria, Australia.
Type locality and stratum:
Locality: Browns Creek outwash section (first gully west of Browns Creek) about 0.4 km east of the mouth of the Johannas River. Aire District, Victoria, Australia.
Stratum: Upper Eocene (Priabonian) carbonaceous clay about 13.1 meters above Notostrea Greensand, Browns Creek Clays (sample E of Cookson and Eisenack 1965).
Stratigraphic and geographic data:
Late Eocene; the species is known only from the type locality, Browns Creek, Victoria, Australia and occurs between 1.8 meters below to 13.1 meters above the Notostrea Greensand.
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