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Andalusiella mauthei ssp. punctata
A. mauthei subsp. punctata, (Jain and Millepied, 1973), Masure, Tea and Yao 1996; Emendation as Andalusiella mauthei subsp. punctata: Masure et al., 1996, p.180-181
Basionym: Palaeocystodinium punctatum Jain and Millepied, 1973
Holotype: Jain and Millepied 1973, pl. 3 fig. 27
Age: Maastrichtian
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Original diagnosis: [Jain and Millepied 1973, p. 29]:
Diagnosis:
Shell spindle-shaped, dorso-ventrally flattened, with two short appendages, one apical and one antapical; Antapical horn divided into a another short horn. Archeopyle trapezoidal below apical horn; periphragm micropunctate. Capsule large, filling completely periphragm cavity, endophragm smooth. Apical horn terminate into a dentate end on free side.
No furrows seen.
Archeopyle not observed in all specimens.
Another thin membrane exterior to periphragm is present, seen only around horns.
The size ratio between horn and central vesicle (H-V ratio) remains mostly 1:4.
Dimensions:
Holotype: Shell length 144 µm, breadth 65 µm; Capsule length 91 µm, breadth 65 µm; Apical horn-length 22.1 µm, breadth 9.1 µm; Antapical horns-length 31.2 µm, breadth 7.8 µm.
Observed range: Shell length 140-150 µm, breadth 55-70 µm; Capsule length 95-97 µm, breadth 60-70 µm; Apical horn-length 22-23 µm, breadth 8-11 µm; Antapical horns-length 9-32 µm, breadth 4-8 µm.
Affinities/Comparison:
Palaeocystodinium punctatum sp. nov. compares only with P. benjaminii Drugg (1967) in having two antapical horns but differs readily in having no fiber or hairs on the apical horn and having dentate apex, micropunctate periphragm and third exterior layer.
Remarks:
The presence of two closely placed antapical horns and horn-vesicle ratio makes it difficult to place this species in Palaeocystodinium and the former evidence might permit the transfer of these forms under a new genus.
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Emended diagnosis:
Masure, Tea and Yao, 1996, p. 180-181:
Diagnosis:
Proximate acrocavate dinoflagellate cyst with rhomboidal to ovoidal central body, one apical horn and two antapical horns associated with adjacent bases, forking close to the central body.
Wall composed of a smooth periphragm and a smooth and brown endophragm with thickenings in horn area. The periphragm on distal end of horns is punctated.
Peridinoid paratabulation indicated by the archeopyle, paracingulum poorly developed, parasulcus with bean-shaped flagellar scar.
Intercalary archeopyle of I/I type, standard 2a hexa, isodeltaforme, operculum free, occasionally adherent.
Remark:
Schrank, 1987 (p. 265) retained this taxon as a subspecies, A. polymorpha subsp. punctata. The subspecies punctata is more related to the species mauthei (Fig. 5), the antapical horns are associated close to the central body and not dissociated as A. polymorpha. A. mauthei and A. mauthei subsp. punctata are acrocavate. The paracingulum of A. mauthei is bordered by rows of small tubercles, but of A. mauthei subsp. punctata it is not.
The two antapical horns of A. mauthei subp. Inflata are punctated, these of A. mauthei are not.
Basionym: Palaeocystodinium punctatum Jain and Millepied, 1973
Holotype: Jain and Millepied 1973, pl. 3 fig. 27
Age: Maastrichtian
-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
Original diagnosis: [Jain and Millepied 1973, p. 29]:
Diagnosis:
Shell spindle-shaped, dorso-ventrally flattened, with two short appendages, one apical and one antapical; Antapical horn divided into a another short horn. Archeopyle trapezoidal below apical horn; periphragm micropunctate. Capsule large, filling completely periphragm cavity, endophragm smooth. Apical horn terminate into a dentate end on free side.
No furrows seen.
Archeopyle not observed in all specimens.
Another thin membrane exterior to periphragm is present, seen only around horns.
The size ratio between horn and central vesicle (H-V ratio) remains mostly 1:4.
Dimensions:
Holotype: Shell length 144 µm, breadth 65 µm; Capsule length 91 µm, breadth 65 µm; Apical horn-length 22.1 µm, breadth 9.1 µm; Antapical horns-length 31.2 µm, breadth 7.8 µm.
Observed range: Shell length 140-150 µm, breadth 55-70 µm; Capsule length 95-97 µm, breadth 60-70 µm; Apical horn-length 22-23 µm, breadth 8-11 µm; Antapical horns-length 9-32 µm, breadth 4-8 µm.
Affinities/Comparison:
Palaeocystodinium punctatum sp. nov. compares only with P. benjaminii Drugg (1967) in having two antapical horns but differs readily in having no fiber or hairs on the apical horn and having dentate apex, micropunctate periphragm and third exterior layer.
Remarks:
The presence of two closely placed antapical horns and horn-vesicle ratio makes it difficult to place this species in Palaeocystodinium and the former evidence might permit the transfer of these forms under a new genus.
-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
Emended diagnosis:
Masure, Tea and Yao, 1996, p. 180-181:
Diagnosis:
Proximate acrocavate dinoflagellate cyst with rhomboidal to ovoidal central body, one apical horn and two antapical horns associated with adjacent bases, forking close to the central body.
Wall composed of a smooth periphragm and a smooth and brown endophragm with thickenings in horn area. The periphragm on distal end of horns is punctated.
Peridinoid paratabulation indicated by the archeopyle, paracingulum poorly developed, parasulcus with bean-shaped flagellar scar.
Intercalary archeopyle of I/I type, standard 2a hexa, isodeltaforme, operculum free, occasionally adherent.
Remark:
Schrank, 1987 (p. 265) retained this taxon as a subspecies, A. polymorpha subsp. punctata. The subspecies punctata is more related to the species mauthei (Fig. 5), the antapical horns are associated close to the central body and not dissociated as A. polymorpha. A. mauthei and A. mauthei subsp. punctata are acrocavate. The paracingulum of A. mauthei is bordered by rows of small tubercles, but of A. mauthei subsp. punctata it is not.
The two antapical horns of A. mauthei subp. Inflata are punctated, these of A. mauthei are not.