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Brigantedinium grande

From Fensome et al., 2019:
Brigantedinium grande Matsuoka, 1987, p.55–56, pl.5, figs.1–2; text-fig.4D ex Head, 1996b, p.1231. Holotype: Matsuoka, 1987,
pl.5, figs.1–2. This name was not validly published in Matsuoka (1987, p.55–56) since the generic name
Brigantedinium was not validly published at that time. Age: Holocene.

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Original description: [Matsuoka, 1987]:

Brigantedinium grande sp. nov.
Plate 5, figure 1-2 ; Text-fig. 4D
1976b Peridinium sp. cf. P. punctulatum Paulsen ; Matsuoka, p. 361-362, pl. Ill, fig. ll.

Derivation of name ; Latin grandis, large ; with reference to the large cyst body.
Diagnosis : Large to intermediate chorate cyst brownish-pigmented, consisting of smooth autophragm, without any ornament on surface. Except for archeopyle, no feature indicating paratabulation. Archeopyle intercalary, broadly hexagonal (lati-thetaform), formed by loss of the 2a paraplate. Archeopyle ratio : 0.6-0.7.
Testa quae est globularis et grandis vel intermedia in magnitudine et brunnea in colore consistit ex levi autophragmate sine ullo ornamento
in superficie. Archeopyla excepta nullus vultus indicat paratabulationem. Archeopyla quae formatur remotione secundae anterioris et intercalaris paraplattae est late hexagonalis.

Holotype : Slide no. SR8-2 (15.2/143.0), PI. 5, figs. 1-2, sample no. SR8, Recent sediment in Lake Saroma, Hokkaido, North Japan.

Dimensions : Cyst diameter 59.7-66.0 μm (Holotype 60.0 μm).
Number of specimens observed : 8.

Remarks : This spherical cyst is different from Bngantedi柁Ium canacoense (Wall) in possessing larger cyst body and a broadly hexagonal archeopyle. This species is also similar to B. majusculum in its larger cyst body, but differs from the latter in bearing a lati-thetaform (hexagonal) archeoyle.

Geographical distribution around the Japanese Islands : Off Hachinohe in North Japan by Matsuoka (1976).

Equivalent thecate form : Probably a species of the subgenus Archaeperidlnium in the genus Protoperidinium based on its hexagonal archeopyle, but the thecate species corresponding to this cyst has been unknown.
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