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Cerodinium kangiliense

Cerodinium kangiliense, Hansen in Nøhr-Hansen and Heilmann-Clausen, 2001, p.158,160,162-164, fig.4, nos.1-9, fig.5, nos.1-9.

Nøhr-Hansen and Heilmann-Clausen (2001, p.158) cited the authorship as "n. sp. ex Hansen 1980" in reference to an unpublished thesis by Hansen. Since the use of this name by Nøhr-Hansen and Heilmann-Clausen (2001) represents its first effective publication as well as its first valid publication, the authorship citation is here given as "Hansen in Nøhr-Hansen and Heilmann-Clausen" rather than "Hansen ex Nøhr-Hansen and Heilmann-Clausen".

Holotype: Nøhr-Hansen and Heilmann-Clausen, 2001, fig.4, nos.1-3.
Paratypes: Nøhr-Hansen and Heilmann-Clausen, 2001, fig. 4 (4-5),
Type locality: Kangilia section (Figs. 1, 3) on the north coast of Nuussuaq, West Greenland GGU sample 210 506. The species is recorded from 715-787 m above sea level; from the middle and upper part of the Eqalulik Formation (Dam pers. Comm. 2000).
Range: first occurrence in the uppermost part of the middle Danian, upper NP 3 Zone; range up into the lowermost Selandian, lower NP 5 Zone
Age: mid Danian-earliest Selandian.

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Original description: [Nøhr-Hansen and Heilmann-Clausen, 2001, p.158,160,162-164]:

Description:
Cyst type: Bivacate, peridinioid.
Shape: Dorso-ventrally compressed cyst, pericyst elongate with a moderately long tapering apical horn and two slightly shorter antapical horns. Endocyst pentagonal to sub-circular.
Wall relationship: The cyst is two-layered, bicavate to cornucavate, composed of a smooth pericyst with short parasutural rows and/or intratabular clusters of granulae or echinae. The endocyst is darker and without and visible ornamentation.
Tabulation: Paratabulation is reflected partly by the archeopyle and by parasutural granulae or short echinae and by intratabular arrangements of granulae or echinae. Peridiniacean, with the formula 4’,3a, 7’’, Xc, 5’’’, 2’’’’, 0-5 s.
Periarcheopyle: Intercalary (2a) large elongate hexagonal periarcheopyle of Cerodinium-type.

Dimensions:
Holotype: overall length 104 μm, width 62 μm, length of apical horn 24 μm, length of antapical horn 24 μm.
Minimum average and maximum dimensions of specimens from Nuusuaq: overall length 90 (111.6) 115 μm, width 54 (59.7) 66 μm, length of apical horn 16 (27.4) 38 μm, length of antapical horn 16 (20) 26 μm.
Minimum, average and maximum dimensions of 9 specimens in from Denmark: Overall length 82 (93.1) 100 μm, width 48 (51.4) 54 μm, length of apical horn 21 (24) 33 μm, length of antapical horn 15 (16.6) 21 μm.

Affinities/Comparison:
Cerodinium kangiliense differs from the somewhat similar species C. dartmoorium Cookson & Eisenack 1965a and C. medcalfii Stover 1973 by being smaller and by its more prominent sculpture elements. C. medcalfii is recorded from the Thanetian of Denmark (Heilmann-Clausen 1985) ans southern England (Powel et al., 1996) and there is no stratigraphical overlap between C. medcalfii and C. kangiliense.
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