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Circulodinium latoaculeum

From Fensome et al., 2019:
Circulodinium latoaculeum (Yun Hyesu, 1981, p.42–43, pl.11, figs.17–19) Prince et al., 1999, p.160. Holotype: Yun Hyesu, 1981, pl.11, fig.18; Fensome et al., 1991, fig.2 — p.657; fig.2 — p.693 (mislabelled as Cleistosphaeridium multifurcatum subsp. multifurcatum); Fensome et al., 1993a, fig.2 — p.1261; Fauconnier and Masure, 2004, pl.35, figs.9–10; Fensome et al., 2019a, fig.20B. NOW Cleistosphaeridium latoaculeum. Originally Cleistosphaeridium multifurcatum subsp. latoaculeum, subsequently Heterosphaeridium latoaculeum, thirdly Circulodinium latoaculeum, fourthly (and now) Cleistosphaeridium latoaculeum. Age: early Santonian.

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Original description: [Yun Hyesu, 1981] Translation: pkb 2024
Material: approx. 25.
Dimensions: max. size of the ZK = 60(65)70, max. length of the spines = 12(14)16, medial width of the spines. Holotype: plate 11, fig. 18. Width of the ZK = 66, length of the spines = up to 12, medial width of the spines = 2-6. Paratype Plate 11, Fig 19.
Diagnosis: A subspecies of C. multifurcatum with broad, flattened satchels that are broadened proximally and distally.
Description: The cyst has a partially granulated layer from which the solid spines form. They are broadened distally and capped at one end, with the distal edge being serrated and sometimes slit. In contrast to the relatively constant length of the spines, the width varies considerably. Wide spines can appear together with narrow ones. In the narrow spines the smallest width is just before the distal end, while in the broad spines it is medial. Relatively larger or longer spines are developed on the poles.
Comparison: The subspecies differs from C. multifurcatum multifurcatum in its predominantly broad, smooth and very uniform spines.
Proven so far: First description, Santon, Germany.
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