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Imperfectodinium bulbosum

Imperfectodinium "*bulbosum" Zevenboom and Santarelli in Zevenboom, 1995, p.147, pl.5, figs.10-12; pl.6, figs.1-4.

Name not validly published: considered a manuscript name. This name was also not validly published in Gedl (1996, p.198), since no description was provided.
Holotype: Zevenboom, 1995, pl.6, figs.1-4.
Age: latest Early Miocene-Late Miocene.

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Original description (Zevenboom, 1995)
Imperfectodinium bulbosum Zevenboom and Santarelli sp.nov. (manuscript name) (Plate VI, 5-10)
1989 Gen. et sp. indet. Piasecki, 1980 Manum et al., plate 10, figs. 1-2

Manuscript holotype: Sample H 280/281, Sg. H 22, Plate VI, 1-4.
Manuscript paratype: Sample H 280/281, Sg. H 24, Plate V, 10-12.
Etymology: in reference to possession of bulbose processes
Type locality: Heumensoord well, The Netherlands
Type stratum: Eibergen Formation

Diagnosis: A relatively small species of Imperfectodinium, characterized by its variable surface ornamentation consisting of low septa and/or crests with hollow bulbose processes, and solid or hollow captitate processes.

Dimensions: Height 35 (38) 45 µm; Width 27 (34) 40 µm; Height of all types of processes 5 (7.5) 10 µm; Wall thickness 1-2.5 µm. N = 30

Description: A species of Imperfectodinium, characterized by its relatively small size and variable surface ornamentation, typically includingmbulbose processes. Endo- and periphargm are closely appressed. The periphragm is faintly microgranulate to microreticulate and gives rise to crests, septae, slender hollow or solid capitate processes and hollow bulbose processes. The ornamentation completely reflects a gonyaulacoid paratabulation. Accesory septae or crests may occur, most notably on the ventral side. The “intraplate” archeopyle is formed by the loss of paraplate 3’’. The archeopyle margin is thickened and serrate.

Discussion: Imperfectodnium bulbosum differs from Imperfectodinium septatum Zevenboom and Santarelli sp. nov. in displaying more than one type of ornamentation.
Occurrences: The Netherlands (this study), Italy (Zevenboom et al., in prep.) and the Norwegian Sea (Manum et al., 1989).
Total stratigraphic range: Latest to early Miocene to Upper Miocene (this study and Manum et al., 1989,, see Fig. 5)

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