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Protobatioladinium mercieri

Protobatioladinium mercieri Feist-Burkhardt and Pross, 1999, p.107–108, pl.1, figs.1–6; pl.2, figs.1–6; pl.7, figs.1–6; pl.11, figs.1–9; pl.12, figs.1–9; text-fig.1, nos.1a–b,2; text-fig.2, nos.1a–b,2a–b,3a–b.

Holotype: Feist-Burkhardt and Pross, 1999, pl.1, figs.1–6; pl.7, figs.1–3; pl.11, figs.1–6; text-fig.1, no.1.
Age: late Bajocian–early Bathonian.

Original description (Feist-Burkhardt & Pross, 1999):
Protobatioladinium mercieri sp.nov. (Pl. 1, fig. 1-6, etc.)
1992 Protobatioladinium sp. – Feist-Burkhardth & Wille, Pl. 2, figs. 3-7
1998 Protobatioladinium sp. 1 – Feist-Burkhardt & MOnteil, Pl. 5, figs. 4, 5.
Derivation of name: In honour of J. Mercier, who was a student of G. Deflandre and did the first palynological work on the Bajocian and Bathonian strata in Calvados where the type material comes from (Mercier, 1938).
Holotype locality and age: Upper part of Calcaire à spingiaires Formation, Port-en-Bessin (West), Normandy, France, Late Bajocian, Parkinsoni Zone, Bomfordi Subzone. For more information see Feist-Burkhardt (1998).

Description: Cyst proximate, cavate or acavate, ovoidal with a long hollow apical horn. Antapex slightly angular rounded, without distinct antapical horn protrusion. The autophragm is thick, frothy porous and may be covered be a second, very thin, translucent outer membrane. Autophragm ornamented irregularly by grana, verrucae and gemmae. Archeopyle compound, type [2A]2I. Archeopyle formed by the loss of three opercular pieces, two intercalary paraplates 1a and 2a, and the apical cap often remains in situ. Paratabulation expressed by archeopyle margin and outline of the opercular pieces only. Paracingulam and parasulcus not indicted.

Size: Holotype: Length x breadth (laterally compressed, apical cap in situ): 93 x 57 µm.
Size range: Length (including apical cap): 64 to 93 µm, breadth: 39 to 57 µm

Remarks: the outline of the opercular pieces and the archeopyle margin suggest a paratabulation with three apical homogenous, homologue ‘1’ in exsert position not reaching the morphological apex and the homologues ‘2’ and ‘3’ forming the apical cap, two intercalary paraplates (1a and 2a) and six precingular paraplates (1’to 6’). This is in accordance with the interpretation of Fensome et al., 1993, p. 77) for the family Pareodiniaceae based inter alia on SEM observations by Below (1990).
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