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Phallocysta erregulensis

Phallocysta ?erregulensis (Filatoff, 1975, p.90, pl.29, figs.12–14) Stover and Helby, 1987a, p.111. Emendation: Stover and Helby, 1987a, p.111, as Phallocysta erregulensis, as a revised description. Holotype: Filatoff, 1975, pl.29, fig.14; Stover and Helby, 1987a, figs.11L–M.

Originally Evansia, subsequently Evansia?, thirdly Phallocysta, fourthly Andreedinium, fifthly (and now) Phallocysta?. This species was retained in Phallocysta by Riding (1994, p.14). Questionable assignment: Riding (1994, p.14). Age: Bajocian

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G.L. Williams short notes on species, Mesozoic-Cenozoic dinocyst course, Urbino, Italy, May 17-22, 1999 - LPP VIEWER CD-ROM 99.5.

Phallocysta erregulensis (Filatoff, 1975) Stover and Helby, 1987a, emend. Stover and Helby, 1987a. According to Stover and Helby (1987a), Phallocystas erregulensis has a type I archeopyle in both the pericyst and endocyst. The perioperculum is linteloid, the endoperculum is rhomboidal and the width is greater than the length. It differs from Phallocysta eumekes in having a more variable shape and in the nature of the archeopyle. In P. erregulensis both the peri- and endoarcheopyle are I; in P. eumekes the periarcheopyle is 3I, the endoarcheopyle has not been delineated. Size: diameter of body 31-36 µm, horn length 19-22 µm, width of horn 17-22 µm.
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Revised description: Stover and Helby, 1987, p. 111
Cysts proximate, generally epicavate, less commonly bicavate, and on rare specimens, the hypocyst may be cornucavate. Pericyst ovoid to trilobate in dorsoventral view, with the apical end being consistently narrower than the antapical end, and except for rare, strongly trilobate forms, cysts longer than wide.
Endocyst subcircular to roundly triangular, occupying practically all of the hypocyst as well as part of the epicyst, and widely separated from the periphragm apically. Epi-pericoel prominent. On some specimens, narrow to relatively wide hypopericoels present (Fig.lOA-D) and these may be confined to the antapical corners (Fig.lOE). Periphragm smooth to uniformly granulate. Endophragm smooth, granulate or with low (up to 1.51 µm long) processes, which may be pointed or capitate (Fig.llD,M). Processes 1-2 µm apart. Archeopyle intercalary, type l; end- and periopercula normally attached adcingularly (Fig.lOB), although the latter may be separated completely on some specimens. Periarcheopyle significantly longer than end-archeopyle (Fig. IOA). Paracingulum not indicated, or expressed by transverse folds in the periphragm. Parasulcus not delimited.
Variability: The greatest variability is in the shape of the cysts and in the amount of cavation (Figs IOA,C-E, I 1).
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Remark Riding 1994, p. 16:
This species is considered to be questionably attributed to Phallocysta due to having an anterior intercalary, type Ia, periarcheopyle and endoarcheopyle according to Stover and Helby (1987, p. 111). Below (1987) stated that Phallocysta has a 3I archeopyle in his emendation of the genus. The transfer of this species to Below is therefore rejected.
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Affinities:
Stover and Helby, 1987, p. 111: Phallocysta erregulensis closely resembles P. eumekes Dörhöfer & Davies emend. Riding 1984 but is distinguished by its more
variable shape and the nature of the archeopyle. In P. erregulensis both the end- and periarcheopyle are type 1, the perioperculum is linteloid in shape whereas the end-operculum is rhomboidal and its width is greater than its length. In contrast, the peri-archeopyle in P. eumekes varies from type I to type 3I. The end-archeopyle
of P. eumekes has not been discussed.
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