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Thalassiphora papulovii

From Fensome et al., 2019:
Thalassiphora papulovii Vasilyeva in Vasilyeva et al., 2001, p.68, pl., figs.3–5,7 ex Vasilyeva in Andreeva-Grigorovich et al., 2011, p.33–34, pl.4, figs.13–16.
Holotype: Vasilyeva et al., 2001, pl.2, fig.3; Andreeva-Grigorovich et al., 2011, pl.4, fig.13.
This species was not validly published in Vasilyeva et al. (2001), since no English or Latin diagnosis was provided.
Age: Late Eocene–Early Oligocene.

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Original description: [Vasilyeva in Vasilyeva et al., 2001, ex Vasilyeva in Andreeva-Grigorovich et al., 2011] (translated from Russian):

Thalassiphora papulovi Vassilyeva sp. nov. Tab. IV, 13-16.

Name. In honor of Professor Georgy Nikolaevich Papulov, a famous geologist.

Diagnosis. Cavate subsphaerical figured cyst of large size. Epicyst and hipocyst with relief outline. Endocyst subsphaerical, oval. Pericel wide. Periphragm dense, smooth, not perforate. Periarcheopyle semicircular, precingular type. Paracingulum not indicated. Paratabulation expressed by periarcheopile.

Description. Cavate cyst is broadly oval, shaped, and large in size. The epicista has complex relief contours in the form of protrusions, protrusions of the periphragm: one central, pointed in shape, narrower, higher, and two symmetrical on the lateral sides of the epicista, wider and rounded. A small wide carina is developed in the antapical part of the pericyst. The lateral sides of the hypocyst have “shoulders” in the form of thickened pointed protrusions, the corners of the periphragm. The endocyst is round, sometimes oval. The pericel is wide. The periphragm is dense, solid, transparent, slightly granular. The endophragm is thin and smooth. The periarchaeopile is precingular, large, semicircular in shape. The paracingulum is practically not formed.

Holotype. Table IV, 13. Rev. 7015 K-1658-6, IGIG Ural Branch RAS.

Location. Kachar quarry, well. 1658, arr. 6, upper part of the Chegan Formation (Kostanay region, Rudny).

Dimensions (µm). Holotype: pericyst length 144; pericyst width-68; the larger and smaller diameters of the endocyst are 66 and 64, respectively; keel height - 7. Other specimens (8 specimens): length and width of pericyst - 155-142; endocyst diameter 89-68; keel height 6-9.

Variability. The morphotype is quite stable and easy to diagnose; The pericyst in outline resembles the figured leaf blade of a plant. There is variability in the outgrowths on the pericyst and displacement of individual corners to the dorsal or ventral side of the pericyst.
In some specimens, a large hole is visible on the hypocyst.

Spreading. Kazakhstan, Southern Trans-Urals, northern part of Turgais. the trough, the roof of the Chegan formation, the Kurgan layers. Upper Eocene, Lower Oligocene.
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