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Vallodinium heilmannii

From Fensome et al., 2019:
Vallodinium heilmannii Iakovleva, 2016, p.329 (p.19 on PDF initially published online), pl.6, figs.3–8; text-fig.4b (part).
Holotype: Iakovleva, 2016, pl.6, fig.3.
Age: earliest Eocene.

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Original description (Iakovleva, 2016):
Vallodinium heilmannii sp. nov. Plate 6, figures 3 8; Figure 4b
Holotype. Plate 6, figure 3; slide MPA 62304-2; England Finder reference P18/2.
Paratype 1. Plate 6, figures 5 6; slide 58145-Ailly-92A19; England Finder reference E34/2.
Paratype 2. Plate 6, figure 4; slide 67642; England Finder reference V38/4.
Type stratum and locality. Sample SMA 2-53, »13.0 m depth; Sotteville-sur-Mer Member; BRGM Scientific Borehole SiegeMadame-2, Dieppe-Hampshire Basin, France.

Diagnosis. A circumcavate species of Vallodinium with a rounded-pentagonal ambitus and ornamentation consisting solely of low sutural folds.

Description. A thin-walled, mostly circumcavate, mid-size cyst with rounded-pentagonal ambitus. Apical and lateral horns are very short; the antapical horns are short and sometimes may be of almost equal length. Periphragm is ornamented by narrow sutural folds, which may be slightly jagged. Endocyst subcircular; endophragm smooth. Endophragm and periphragm may be appressed over parts of the cyst. Archaeopyle equiepeliform; operculum free, in place or partially attached.

Dimensions. Holotype: pericyst length 84 µm; pericyst width 76 µm; endocyst length 68 µm; endocyst width 63 µm. Paratype 1: pericyst length 78 µm; pericyst width 80 µm; endocyst length 63 µm; endocyst width 65 µm. Paratype 2: pericyst length 86 µm; pericyst width 78 µm; endocyst length 71 µm; endocyst width 68 µm.

Dimensions of measured specimens: pericyst length 63-88 µm (mean value 74 µm); pericyst width 60-87 µm (mean value 74 µm); endocyst length 48-73 µm (mean value 60.5 µm); endocyst width 51-75 µm (mean value 64 µm). Twenty-seven specimens measured.

Comparison. Vallodinium heilmannii differs from other species of Vallodinium by narrow sutural folds, poorly developed horns, rounded-pentagonal shape and its smaller size.

Stratigraphical range. Earliest Eocene.
Derivation of name. In honor of the Danish palynologist Claus Heilmann-Clausen.
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