Back
Baserus swabiana
Baserus swabiana, (Morbey, 1975, p.39–40, pl.14, figs.5–11; pl.17, figs.4–9; text-figs.12a–c,13a–c,14a–b,15), Özdikmen, 2009, p.237. Emendation: Below, 1987a, p.94–96 (as Suessia swabiana).
NOW Suessia. Originally (and now) Suessia, subsequently Baserus (generic name illegitimate). Below (1987a, p.94) considered Rhombodella (now Heibergella) kendelbachia to be a questionable junior synonym of this species.
Holotype: Morbey, 1975, pl.14, fig.5; pl.17, fig.4; text-figs.12a–c; Fensome et al., 1995, figs.1,7–9 — p.1819.
Age: Rhaetian.
-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
Original description: [Morbey, 1975, p. 39-40]:
Diagnosis:
Cyst spheroidal, ovoidal, spherical; proximate, pseudocavate, tabulate, compression dominantly dorso-ventral.
Wall thin, membranous, microgranulate, microreticulate, granulate, rarely sculptured with fine spinae in part. Periphragm closely appressed to or apically/equatorially partially detached, but not completely detached, from endophragm.
Cingulum laevorotatory, defined occasionally by secondary folds. Secondary folding usually peripheral and along length of cyst.
Sulcus: exact nature indiscernible, defined and circumscribed by sulcal "plates".
Archeopyle: a combination of apical and anterior intercalary "plates".
Tabulation complex, defined by a visual coarse reticulation of plate-bounding aseptate sutures, which barely modify cyst outline. Approximately ?4 apical (?4"), ?3-6 anterior intercalary (?3-6a), 7-9 postapical (7-9ap), 12-14 precingular (12-14 ), 12-14 cingular (12-14c), 12-14 postcingular (12-14" ), 7-9 posterior intercalary (7-9p), 6 preantapical (6pa), 1 antapical (1""), and 4-8 sulcal "plates" (4-8S). The epitractal sutures are often poorly represented.
Dimensons:
Apical length/diameter (periphragm) 42 (49) 65 µm (31 specimens measured).
Apical length/diameter (endophragm) 40 (46) 52 µm. Cingulum diameter (periphragm) 33 (41) 52 µm.
-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
Emended diagnosis:
Below, 1987, p. 95-96:
Diagnosis:
Arrangement of vesicles / plates on the dinoflagellate amphiesma cop, pop, cap, 5", 3a-4a, 7"", 7c, 5"""-6""", 2"""",as, ns, ps; as usually connected dextrally with 5" and 7""; partiform arrangement of 1"""" and 2"""", 1"""" borders on ps and the central sulcus, 2"""" is pentagonal or hexagonal.
Cystic habit proximate, ovoidal, biconical, with a coniform epicyst and obtusi-coniform hypocyst, apex acute, antapex flattened, indented, and then with its pole located sinistrally, equatorial diameter circular to oval; size large; surface glabrous, divided into areae by finate, low to broad muri, muri distally unornamented or covered unevenly with very fine granules, rows of granules partially fused, the muri can also take the form of low, narrow, distally smooth or scabrate ribs; areation multiareate, multiserial; areation formula n[3], n[2], n[a], n[1], nc, n(1), n(2), n(3), n(4), sas, pas, NR rs/Xrs, NR ls/ls, pps, sps; number and shape of areae individually variable, between 70 and 100* (* numbers exemplified in the present material), small tetragonal to octagonal areae distributed as follows: apical 13 and smaller 23; multiareate secanterior series n[2] (5-6)* comprising large, iso- to steno- delta-/theta-form pentagonal to hextagonal areae and asymmetrical polyhedra; (7-9)* large iso- to steno-, delta- but also theta-form pentagonal, hexaxonal, and heptagonal areae in the anterior intercalary series, which begins on the dextral side of area column 1[1] + 1[2]; multiareate primanterior series n[1] (12-16)* with small, planate, pentagonal, hexagonal, seldom heptagonal areae, always beginning with a larye iso/steno heptagonal 1[1] and ending with a broad, oblique area n(1), areae frequently with alternating geniculate or linear contact with the anterior intercalaries; multiareate cingulum comprising (13-16)* small, usually iso hexagonal, seldom planate or pentagonal areae; primposterior series n(1) with (13-19)* small, iso pentagonal and hexagonal, seldom planate areae; secposterior series with n(2) (9-14)* iso pentagonal and hexagonal, also heptagonal and octagonal areae; (7-10)* large, iso/steno pentagonal and hexagonal tertposterior, areae n(3), reduced in size from dorsal to ventral, the area column 1(1) + 1(2) + 1(3) usually displaced somewhat to the posterior in relation to the next column sinistrad; antapex consisting of three iso pentagonal and hexagonal, also heptagonal, areae of the quartposterior series; hypocystal intercalary area in variable position; cingulum equatorial, cingulum and area series helicoid laevorotatory and displaced at the sulcus, multiareate, sulcus slightly indented, sas usually typically steno or iso omegaform-pentagonal, pas posteriorly non-finate, oriented obliquely to the cingulum anteriorly to the last primanterior area, rs and ls not reflected separately, pps and sps Iarge, polyhedral; disintegration archaeopyle, tn[3] + tn[2] + n[a], comprising apical, sinistral and dorsal, and seldom dextrolateral, regions anterior to the primanterior series, but always without the last three or four dextral areas of the anterior intercalary series and sas, operculum solvate, dividing into individual pieces or groups, general opercular formula 1[3]s + 2[3]a + 1[2]s + 2[2]s + 3[2]s + n[2]s + 1[a]s + 2[a]s + 3[a]s + 4[a]s + n[a]s. Pellicular habit proximate, ovaloidal, peridinoidal; surface unornamented, smooth, scabrate, in some places allowing traces of the areation of the enclosed cyst to show through, or partially tracing it with low ribs, or divided into areae by means of low ridges or low septae; aration pattern NR PR/XPR, NR"/5", NRa/3a-4a, NR""/7"", NRc/Xc/7c, NR"""/5"""-6""", NR""""/2"""", NRs/Xs, as, ns, ps, PR undifferentiated; steno V 1"(V), VI 2"(VL), deltaform inverse-camerate 3" (DDL), deltaform inverse-camerate 4" (DR), VII 5"(VR); deltaform inverse-camerate 1a(LDL), deltaform linteloid 2a (D), deltaform inverse-camerate 3a (R); anteriorly geniculate V-nE 1"" (VVL), anteriorly geniculate V-nE 3"" (DL), anteriorly linear IV-nE 4"" (D), anteriorly geniculate V-nE 5"" (DR), anteriorly geniculate V-nE 6"" (RVR), anteriorly linear IV-nE 7"" (VVR); or PR undifferentiated, V 1", (V), VI 2" (VL), VI 3" (L), V 4" (R), V 5" (VR), deltaform inverse-camerate 1a (L), deltaform inverse-camerate 2a (DL), deltaform fastigiate 3a (D), deltaform inverse-camerate 4a; VII-nE 1" (V), V-nE 5" (VR), anteriorly geniculate V-nE 7"" (VVR); cingulum helicoid laevorotaory, nonareate or partite into 7 lati pentagonal and lati hexagonal areae, displaced at the sulcus, borders of the cingulum formed by low crests or septa; 6 postcingulars, camerate 1""" (VVL), posteriorly linear IV-nE 2""" (VL), posteriorly geniculate V-nE 3""" (DL), posteriorly linear IV-nE (D), posteriorly linear IV-nE 5"""(DR), posteriorly geniculate V-nE 6""" (VR); omegaform linteloid 1"""" (LVL), omegaform linteloid 2"""" (CD); or only 5 postcingulars, camerate 1""" (VVL), posteriorly linear IV-nE 2""" (VL), posteriorly geniculate V-nE 3""" (DL), posteriorly linear IV-nE 4""" (DR) and posteriorly geniculate V-nE 5""" (VR); sulcus broad and shallow, undivided, anterior end dextrally in contact with 5" and 7"" or only with 7""; pylome opened by irregular thinning, bursting, tearing, or bending away of the pellicle over the cystic archaeopyle.
NOW Suessia. Originally (and now) Suessia, subsequently Baserus (generic name illegitimate). Below (1987a, p.94) considered Rhombodella (now Heibergella) kendelbachia to be a questionable junior synonym of this species.
Holotype: Morbey, 1975, pl.14, fig.5; pl.17, fig.4; text-figs.12a–c; Fensome et al., 1995, figs.1,7–9 — p.1819.
Age: Rhaetian.
-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
Original description: [Morbey, 1975, p. 39-40]:
Diagnosis:
Cyst spheroidal, ovoidal, spherical; proximate, pseudocavate, tabulate, compression dominantly dorso-ventral.
Wall thin, membranous, microgranulate, microreticulate, granulate, rarely sculptured with fine spinae in part. Periphragm closely appressed to or apically/equatorially partially detached, but not completely detached, from endophragm.
Cingulum laevorotatory, defined occasionally by secondary folds. Secondary folding usually peripheral and along length of cyst.
Sulcus: exact nature indiscernible, defined and circumscribed by sulcal "plates".
Archeopyle: a combination of apical and anterior intercalary "plates".
Tabulation complex, defined by a visual coarse reticulation of plate-bounding aseptate sutures, which barely modify cyst outline. Approximately ?4 apical (?4"), ?3-6 anterior intercalary (?3-6a), 7-9 postapical (7-9ap), 12-14 precingular (12-14 ), 12-14 cingular (12-14c), 12-14 postcingular (12-14" ), 7-9 posterior intercalary (7-9p), 6 preantapical (6pa), 1 antapical (1""), and 4-8 sulcal "plates" (4-8S). The epitractal sutures are often poorly represented.
Dimensons:
Apical length/diameter (periphragm) 42 (49) 65 µm (31 specimens measured).
Apical length/diameter (endophragm) 40 (46) 52 µm. Cingulum diameter (periphragm) 33 (41) 52 µm.
-------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------
Emended diagnosis:
Below, 1987, p. 95-96:
Diagnosis:
Arrangement of vesicles / plates on the dinoflagellate amphiesma cop, pop, cap, 5", 3a-4a, 7"", 7c, 5"""-6""", 2"""",as, ns, ps; as usually connected dextrally with 5" and 7""; partiform arrangement of 1"""" and 2"""", 1"""" borders on ps and the central sulcus, 2"""" is pentagonal or hexagonal.
Cystic habit proximate, ovoidal, biconical, with a coniform epicyst and obtusi-coniform hypocyst, apex acute, antapex flattened, indented, and then with its pole located sinistrally, equatorial diameter circular to oval; size large; surface glabrous, divided into areae by finate, low to broad muri, muri distally unornamented or covered unevenly with very fine granules, rows of granules partially fused, the muri can also take the form of low, narrow, distally smooth or scabrate ribs; areation multiareate, multiserial; areation formula n[3], n[2], n[a], n[1], nc, n(1), n(2), n(3), n(4), sas, pas, NR rs/Xrs, NR ls/ls, pps, sps; number and shape of areae individually variable, between 70 and 100* (* numbers exemplified in the present material), small tetragonal to octagonal areae distributed as follows: apical 13 and smaller 23; multiareate secanterior series n[2] (5-6)* comprising large, iso- to steno- delta-/theta-form pentagonal to hextagonal areae and asymmetrical polyhedra; (7-9)* large iso- to steno-, delta- but also theta-form pentagonal, hexaxonal, and heptagonal areae in the anterior intercalary series, which begins on the dextral side of area column 1[1] + 1[2]; multiareate primanterior series n[1] (12-16)* with small, planate, pentagonal, hexagonal, seldom heptagonal areae, always beginning with a larye iso/steno heptagonal 1[1] and ending with a broad, oblique area n(1), areae frequently with alternating geniculate or linear contact with the anterior intercalaries; multiareate cingulum comprising (13-16)* small, usually iso hexagonal, seldom planate or pentagonal areae; primposterior series n(1) with (13-19)* small, iso pentagonal and hexagonal, seldom planate areae; secposterior series with n(2) (9-14)* iso pentagonal and hexagonal, also heptagonal and octagonal areae; (7-10)* large, iso/steno pentagonal and hexagonal tertposterior, areae n(3), reduced in size from dorsal to ventral, the area column 1(1) + 1(2) + 1(3) usually displaced somewhat to the posterior in relation to the next column sinistrad; antapex consisting of three iso pentagonal and hexagonal, also heptagonal, areae of the quartposterior series; hypocystal intercalary area in variable position; cingulum equatorial, cingulum and area series helicoid laevorotatory and displaced at the sulcus, multiareate, sulcus slightly indented, sas usually typically steno or iso omegaform-pentagonal, pas posteriorly non-finate, oriented obliquely to the cingulum anteriorly to the last primanterior area, rs and ls not reflected separately, pps and sps Iarge, polyhedral; disintegration archaeopyle, tn[3] + tn[2] + n[a], comprising apical, sinistral and dorsal, and seldom dextrolateral, regions anterior to the primanterior series, but always without the last three or four dextral areas of the anterior intercalary series and sas, operculum solvate, dividing into individual pieces or groups, general opercular formula 1[3]s + 2[3]a + 1[2]s + 2[2]s + 3[2]s + n[2]s + 1[a]s + 2[a]s + 3[a]s + 4[a]s + n[a]s. Pellicular habit proximate, ovaloidal, peridinoidal; surface unornamented, smooth, scabrate, in some places allowing traces of the areation of the enclosed cyst to show through, or partially tracing it with low ribs, or divided into areae by means of low ridges or low septae; aration pattern NR PR/XPR, NR"/5", NRa/3a-4a, NR""/7"", NRc/Xc/7c, NR"""/5"""-6""", NR""""/2"""", NRs/Xs, as, ns, ps, PR undifferentiated; steno V 1"(V), VI 2"(VL), deltaform inverse-camerate 3" (DDL), deltaform inverse-camerate 4" (DR), VII 5"(VR); deltaform inverse-camerate 1a(LDL), deltaform linteloid 2a (D), deltaform inverse-camerate 3a (R); anteriorly geniculate V-nE 1"" (VVL), anteriorly geniculate V-nE 3"" (DL), anteriorly linear IV-nE 4"" (D), anteriorly geniculate V-nE 5"" (DR), anteriorly geniculate V-nE 6"" (RVR), anteriorly linear IV-nE 7"" (VVR); or PR undifferentiated, V 1", (V), VI 2" (VL), VI 3" (L), V 4" (R), V 5" (VR), deltaform inverse-camerate 1a (L), deltaform inverse-camerate 2a (DL), deltaform fastigiate 3a (D), deltaform inverse-camerate 4a; VII-nE 1" (V), V-nE 5" (VR), anteriorly geniculate V-nE 7"" (VVR); cingulum helicoid laevorotaory, nonareate or partite into 7 lati pentagonal and lati hexagonal areae, displaced at the sulcus, borders of the cingulum formed by low crests or septa; 6 postcingulars, camerate 1""" (VVL), posteriorly linear IV-nE 2""" (VL), posteriorly geniculate V-nE 3""" (DL), posteriorly linear IV-nE (D), posteriorly linear IV-nE 5"""(DR), posteriorly geniculate V-nE 6""" (VR); omegaform linteloid 1"""" (LVL), omegaform linteloid 2"""" (CD); or only 5 postcingulars, camerate 1""" (VVL), posteriorly linear IV-nE 2""" (VL), posteriorly geniculate V-nE 3""" (DL), posteriorly linear IV-nE 4""" (DR) and posteriorly geniculate V-nE 5""" (VR); sulcus broad and shallow, undivided, anterior end dextrally in contact with 5" and 7"" or only with 7""; pylome opened by irregular thinning, bursting, tearing, or bending away of the pellicle over the cystic archaeopyle.