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Coronifera hebospina
From Fensome et al., 2019:
Coronifera hebospina (Yun Hyesu, 1981, p.23–24, pl.12, figs.2,4,6a–b; text-fig.6b) Peyrot, 2011, p.288. Holotype: Yun Hyesu, 1981, pl.12, figs.6a–b; Fensome et al., 1991, figs.3–4 — p.643; fig.3 — p.697. Originally Coronifera oceanica subsp. hebospina, subsequently Coronifera pedata subsp. hebospina, thirdly (and now) Coronifera hebospina. Lentin and Williams (1989, p.77) inadvertently both transferred this taxon to Coronifera pedata and retained it as a subspecies of Coronifera oceanica. Lentin and Williams (1993, p.131) retained this taxon as a subspecies of Coronifera oceanica. Age: early Santonian.
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Original description (Yun Hyesu, 1981) [translation: pkb 2025]
Material: approx. 20 copies.
Dimensions: max diameter of the ZK = 42(45)54, max. length of the spines = 18(19)24, length of the AHH = 18(20)24, number of spines = 85-120
Holotype: Plate 12, Fig. 6, diameter of the ZK = 38x42. Max. length of the spines = 22, length of the AAH = 24, number of spines = approx. 100, width of the paracinglum = 5. Paratype Plate 12, Fig. 4.
Diagnosis: The cyst is oval in outline and tapers slightly apically. It consists of an endophragm and a periphragm, which forms the spines and the AAH. The spines, which taper continuously distally and end in a rounded or ball-shaped manner, are simple or connected to form double spines. The spine bases are arranged roughly in squares on the surface of the CC by ridges; they are arranged in a circle parallel to the paracingulum. The crossbars also bridge the spine-free paracingulum and thus connect two spines on the edges of the paracingulum in pairs. The hollow, distally open AAH is serrated at the free end and has its largest diameter here. The archaeopyle is precingulate.
Description: see diagnosis.
Comparison: The subspecies differs from C. oceanica oceanica except in the smaller number and greater length of the spines due to the simple, short rounded spine ends. They never have hooks whereas bifurcated spines often appear. In C. oceanica oceanica, the spines sometimes have hooks or two needle-shaped appendages that form an obtuse angle and point obliquely upwards or laterally. In addition, a maximum of 5 spines can be connected to each other up to 2/3 of their height by a thin membrane. While the spines in C. oceanica ssp. hebospina continuously taper towards the tip, in O. oceanica oceanica they can have the same diameter proximally and distally. The differences between C. alberti Millioud 1969 include the larger number of finer and longer spines. In terms of the number of spines, C. oceanica hebospina occupies an intermediate position between C. oceanica oceanice and C. alberti.
Coronifera hebospina (Yun Hyesu, 1981, p.23–24, pl.12, figs.2,4,6a–b; text-fig.6b) Peyrot, 2011, p.288. Holotype: Yun Hyesu, 1981, pl.12, figs.6a–b; Fensome et al., 1991, figs.3–4 — p.643; fig.3 — p.697. Originally Coronifera oceanica subsp. hebospina, subsequently Coronifera pedata subsp. hebospina, thirdly (and now) Coronifera hebospina. Lentin and Williams (1989, p.77) inadvertently both transferred this taxon to Coronifera pedata and retained it as a subspecies of Coronifera oceanica. Lentin and Williams (1993, p.131) retained this taxon as a subspecies of Coronifera oceanica. Age: early Santonian.
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Original description (Yun Hyesu, 1981) [translation: pkb 2025]
Material: approx. 20 copies.
Dimensions: max diameter of the ZK = 42(45)54, max. length of the spines = 18(19)24, length of the AHH = 18(20)24, number of spines = 85-120
Holotype: Plate 12, Fig. 6, diameter of the ZK = 38x42. Max. length of the spines = 22, length of the AAH = 24, number of spines = approx. 100, width of the paracinglum = 5. Paratype Plate 12, Fig. 4.
Diagnosis: The cyst is oval in outline and tapers slightly apically. It consists of an endophragm and a periphragm, which forms the spines and the AAH. The spines, which taper continuously distally and end in a rounded or ball-shaped manner, are simple or connected to form double spines. The spine bases are arranged roughly in squares on the surface of the CC by ridges; they are arranged in a circle parallel to the paracingulum. The crossbars also bridge the spine-free paracingulum and thus connect two spines on the edges of the paracingulum in pairs. The hollow, distally open AAH is serrated at the free end and has its largest diameter here. The archaeopyle is precingulate.
Description: see diagnosis.
Comparison: The subspecies differs from C. oceanica oceanica except in the smaller number and greater length of the spines due to the simple, short rounded spine ends. They never have hooks whereas bifurcated spines often appear. In C. oceanica oceanica, the spines sometimes have hooks or two needle-shaped appendages that form an obtuse angle and point obliquely upwards or laterally. In addition, a maximum of 5 spines can be connected to each other up to 2/3 of their height by a thin membrane. While the spines in C. oceanica ssp. hebospina continuously taper towards the tip, in O. oceanica oceanica they can have the same diameter proximally and distally. The differences between C. alberti Millioud 1969 include the larger number of finer and longer spines. In terms of the number of spines, C. oceanica hebospina occupies an intermediate position between C. oceanica oceanice and C. alberti.